Cyathea andina (H.Karst.) Domin (1929: 263)

Lehnert, Marcus, 2011, Species of Cyathea in America related to the western Pacific species C. decurrens, Phytotaxa 26, pp. 39-59 : 53

27. Cyathea andina (H.Karst.) Domin (1929: 263) . Hemitelia andina Karsten (1856: 452) . Type:— COLOMBIA. Magdalena: Santa Marta, 2500 m, Karsten s.n. (holotype B, photo GH).

Hemitelia servitensis Karsten (1856: 451) . Type:— COLOMBIA. “Crescit in silvis humidis umbrosis Novo- Granatae Provinciae Cundinamarcae altitudine 2000 metr. inter Susumuco et Servita,” Karsten s.n. (holotype not located, W?, isotypes “Servita, Bogotá,” B [Herb. Mettenius], P, photo GH).

Hemitelia escuquensis Karsten (1869: 181) . Cyathea escuquensis (H.Karst.) Domin (1929: 263) . Type:— VENEZUELA. “Sinus Maracaibensis prope Escuque, 100 m,” Karsten s.n. (holotype not located, W?, probable isotype “ Escuque, Venezuela,” Karsten 28, B [Herb. Mettenius]).

Hemitelia joadii Baker (1891: 187) . Cyathea joadii (Baker) Domin (1929: 263) . Type:— COLOMBIA. Magdalena: Santa Marta, December 1863, Joad s.n. (holotype K).

Distribution and habitat: — Haiti, Puerto Rico, Colombia, Venezuela, Ecuador, Peru, Bolivia, in humid forests between 400–1900 m.

Remarks: —Plants from the Guianas treated as Cyathea andina belong to a separate species, Cyathea boryana , which is easily distinguished by its pale brown to white petiole scales (vs. dark castaneous with narrow white margins in C. andina , Fig. 2 ).

Figures

FIGURE 2. Trunk morphology. A. Cyathea multiflora, Prov. Pichincha, Ecuador. B–C. Cyathea parvula, Puerto Rico; B. Detail of emerging crosier; C. Detail of persisting spiny petioles on trunk. D. Cyathea leucofolis, Edo. Paraná, Brazil. E. Cyathea boryana, French Guiana. F. Cyathea andina, crosiers only visible after removal of two fronds, Puerto Rico. G–J. Cyathea rufescens, Dept. San Martin, Peru; G. Plant growing on ridge, with several small adventitious plants at the trunk base; H. Detail of frond; J. Trunk apex (A–D, F–J. © Marcus Lehnert, E. © Michel Boudrie).