Diogenidae, , de Saint Laurent and McLaughlin, 2000

McLaughlin, P. A., 2003, Illustrated keys to families and genera of the superfamily Paguroidea (Crustacea: Decapoda: Anomura), with diagnoses of genera of Paguridae, Memoirs of Museum Victoria 60 (1), pp. 111-144 : 113-114

Key to genera of Diogenidae

1. Well developed arthrobranchs present on arthrodial membranes at bases of cheliped and maxilliped 3; pleurobranch present on somite XI (thoracomere 5, above pereopod 2) ( Fig. 4a).......................... 2

— Reduced or vestigial arthrobranchs present on arthrodial membranes at bases of cheliped and maxilliped 3; no pleurobranch present on somite XI (thoracomere 5, above pereopod 2) ( Fig. 4c).......................................... Pseudopaguristes McLaughlin, 2002

2. 14 pairs of gills; pleurobranch present on somite XIV (thoracomere 8, above pereopod 5) ( Fig. 4a)........... 3

— 13 pairs of gills; no pleurobranch present on somite XIV (thoracomere 8, above pereopod 5) ( Fig. 4b)........ 14

3. Endopod of maxillule with well developed external lobe ( Fig. 4k)................................... 4

— Endopod of maxillule without well developed external lobe ( Fig. 4l)................................ 8

4. Ischium of maxilliped 3 with well developed crista dentata ( Figs 2b–f)............................ 5

— Ischium of maxilliped 3 without well developed crista dentata ( Fig. 2a)............................. 12

5. Chelipeds equal or unequal, each with stridulatory mechanism developed on mesial face of palm ( Fig. 6a) 6

— Chelipeds markedly unequal, left largest; neither with stridulatory mechanism developed on mesial face of palm ( Fig. 6b)..... Allodardanus Haig and Provenzano, 1965

6. Chelipeds with acute, corneous-tipped spines on carpi and chelae; males often with pleopod 2 paired, endopod well developed, reduced or absent ( Fig. 7g)..................................... Strigopagurus Forest, 1995

— Chelipeds with tubercles or transverse striate on carpi and chelae; males without pleopod 2 paired............ 7

7. Chelipeds equal or left larger; carpus and palm with transverse striae bordered with fine setae ( Fig. 6c); dactyls of ambulatory legs equal to or longer than propodi; females with unpaired pleopods 2–5 egg-carrying..................................... Ciliopagurus Forest, 1995

— Chelipeds equal, carpus and palm covered with generally blunt tubercles; dactyls of ambulatory legs much shorter than propodi; females with unpaired pleopod 5 non egg-carrying................ Trizopagurus Forest, 1952

8. Chelae symmetrical, together forming operculum ( Fig. 6d); uropods symmetrical ( Fig. 8j).................................. Cancellus H. Milne Edwards, 1836

— Chelae symmetrical or asymmetrical, together not forming operculum; uropods generally asymmetrical........ 9

9. Chelipeds unequal, right distinctly larger.................................. Petrochirus Stimpson, 1858

— Chelipeds subequal or unequal, left usually at least slightly larger............................... 10

10. Shield with prominent Y-shaped linea in posterior half ( Figs 8a, c); posterior carapace primarily membranous; left cheliped slightly to considerably larger than right....11

— Shield without prominent Y-shaped linea ( Fig. 8b, l); posterior carapace well calcified; chelipeds subequal....................... Tisea Morgan and Forest, 1991

11. Rostrum triangular; ocular acicles each with prominent triangular or subtriangular acicular projection ( Figs 8a, b); chelipeds and ambulatory legs with ring-like transverse striae ( Fig. 6e); females with well-developed brood pouch ( Fig. 7d).................... Aniculus Dana, 1852

— Rostrum broadly rounded or obsolete ( Fig. 8c, l); ocular acicles each with subrectangular or subquadrate acicular projection ( Fig. 8c); chelipeds and ambulatory legs most often without ring-like transverse striae; females without brood pouch............... Dardanus Paul’son, 1875

12. Antennal flagella with microscopic setae.................................. Pseudopagurus Forest, 1952

— Antennal flagella with paired, moderate to long setae ( Fig. 2k).................................. 13

13. Chelipeds equal or slightly subequal, similar, dactyls opening in generally horizontal plane ( Fig. 6f).........…......................... Isocheles Stimpson, 1858

— Chelipeds unequal and dissimilar; dactyls opening in almost vertical plane ( Fig. 6g). Loxopagurus Forest, 1964

14. Males with pleopods 1 and/or 2 paired, modified as gonopods ( Figs 7a, e, f); females with ( Figs 2m, 7b, c) or without pleopod 1 paired, modified.............. 15

— No paired pleopods in either sex................ 16

15. Pereopod 4 chelate ( Fig. 5t); unpaired pleopods 3–5 occurring on either right or left side of abdomen................................ Paguropsis Henderson, 1888

— Pereopod 4 not chelate; unpaired pleopods 3–5 occurring on left side of abdomen only... Paguristes Dana, 1851

16. Chelipeds subequal ( Fig. 1f).................... 17

— Chelipeds unequal, left appreciable larger ( Figs 1d, e, g).… .......................................... 18

17. Ocular acicles bi or multispinose, contiguous or closely set ( Fig. 1f, 8b); posterior margin of abdominal somite 6 unarmed.................. Clibanarius Dana, 1852

— Ocular acicles simple, widely separated ( Figs 3a, d); posterior margin of abdominal somite 6 spinulose............................. Bathynarius Forest, 1989

18. Rostrum obsolete, roundly subtriangular or broadly round- ed, intercalary rostral process present, well developed, reduced or vestigial ( Fig. 3i)...... Diogenes Dana, 1851

— Rostrum moderate to well developed, triangular, intercalary rostral process absent... Calcinus Dana, 1851