Sanicula orthacantha var. orthacantha

Li, Hui-Min, Zhou, Wei & Song, Chun-Feng, 2022, Taxonomic studies on the genus Sanicula (Apiaceae) from China (): The clarification of some morphological distinction between S. orthacantha var. orthacantha and S. orthacantha var. brevispina, with the reduction of S. petagnioides to the synonymy of the former, and S. orthacantha var. stolonifera to, Phytotaxa 548 (1), pp. 1-25 : 19-23

1a. Sanicula orthacantha var. orthacantha . Figs. 1, 3‒5, 7, 9, 14, 16.

= Sanicula petagnioides Hayata (1908: 103) , syn. nov. Type:— CHINA, Taiwan , Mount Morrison (=Yu Shan), 7500 ped. (= 2286 m), 15 October 1906, T. Kawakami & U. Mori 2026 (TAIF18726!, lectotype designated here; isolectotype TAIF18727!). Remaining syntypes: same locality, October 1906, G. Nakahara s.n. (TI00013635!). Fig. 3.

= Sanicula henryi Wolff (1913: 55) .

Type:— CHINA, Yunnan, Mengtze (= Mengzi), 8000 ft (= 2438 m), A. Henry 10500 [K000697284!, lectotype designated by Pimenov (2017); isolectotypes E00000049!, MO345421!, NY00406257!, US00126973!]. Fig. 4 .

= Sanicula nanchuanensis Shan (1943: 111) .

Type:— CHINA, Chongqing, Nanchuan, King Fu (=Jinfo Shan), Fong-feng-szu (= Fenghuang Temple), 2000 m, 9 June 1935, K.L. Chu 1212 (holotype NAS00028652!; isotypes IBK00338581!, NAS00028653!, PE00754814!, PE00754815!, PE00754816!). Fig. 5A, B .

Diagnostic characters:—Rhizome short, tuberlike, woody, oblique rootstock bearing elongated, fibrous roots, occasionally with fleshy stoloniferous.

Distribution and habitat:— Sanicula orthacantha var. orthacantha is widely distributed in China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Taiwan, Yunnan, Zhejiang), and Vietnam (Fig. 17). It grows on mountain slopes under forest or along ravine, stream at elevations of 600‒2140 m above sea level. It was recorded occur in Cambodia, India, Laos ( Sheh & Phillippe 2005), but we have not as yet seen any material from these countries.

Phenology:—Flowering and fruits from March to October.

Additional specimens examined:— CHINA. Anhui: Chizhou, Anonymous 17681 (NAS), Anonymous 78748 (NAS), Anhui Wild Plant Comprehensive Utilization Committee Exped. 5320 (NAS), R.C. Ching 2684 (NAS); Huangshan, Anonymous 3288 (PE), M.B. Deng & J.Q. Pan 85 (NAS), K. Yao 79038 (NAS); Jingxian, Anonymous 176 (NAS); Ningguo, M.B. Deng et al. 7684 (NAS); Shexian, M. Liu et al. A110011 (KUN, PE), Wild Plant Resource Exped. 1372 (NAS). Chongqing: Chengkou, T.L. Dai 100582 (CDBI, IBK, PE), T.L. Dai 105281 (NAS); Fengjie, H.F. Chou & H.Y. Su 108207 (WUK); Kaizhou, Bashan Exped. 2323 (PE); Nanchuan, H.F. Chang 133 (PE), H.F. Chang 134 (PE), H.F. Chang 216 (PE), S.C. Chen & K.Y. Lang 2399 (PE), W.P. Fang 918 (NAS, PE, US), W.P. Fang 1146 (E, K, LBG, NAS, US), W.P. Fang 1147 (NAS), C.H. Hsiung & T.L. Chou 90548 (PE), C.H. Hsiung & T.L. Chou 90668 (PE), Y.Y. Ho 4640 (NAS), C.H. Hsiung & T.L. Chou 90838 (PE), H.M. Li, Y.S. Zhang & X. Zhang 1140 (NAS), H.M. Li, Y.S. Zhang & X. Zhang 1141 (NAS), H.M. Li, Y.S. Zhang & X. Zhang 1142 (NAS), H.M. Li, Y.S. Zhang & X. Zhang 1144 (NAS), H.M. Li, Y.S. Zhang & X. Zhang 1145 (NAS), K.F. Li 61671 (KUN, NAS), K.F. Li 62310 (KUN, NAS), Plant Geog. Exped. 20 (PE), M.L. Sheh 83659 (NAS), M.L. Sheh 83669 (NAS), Sichuan Exped. 230 (CDBI, PE, SM), T.H. Tu 2917 (IBK, PE), Q.C. Wang 23 (CDBI), L. Wu et al. 4513 (BNU), T. Zhang et al. 10CS1728 (KUN); Pengshui, Anonymous 109 (SM), Anonymous 454 (SM), Anonymous 996 (SM); Qianjiang, Anonymous 429 (SM); Shizhu, Anonymous 668 (SM), Y. Chen 105 (CDBI), Y. Chen 870 (CDBI), Y. Chen 2115 (CDBI), H.M. Li, Y.S. Zhang & X. Zhang 1124 (NAS), W.H. Wang 1337 (CDBI); Wulong, Anonymous 807 (SM); Wuxi, Anonymous 73W-711 (SM), Y.S. Chen et al. 1293 (WUK), P.Y. Li 3510 (NAS), K.H. Yang 58875 (IBK, PE); Youyang, Biol. Depart. Youyang Norm. Sch. 2194 (PE). Fujian: Guangze, Wuyishan Exped. 801248 (FJIDC); Jiangle, Longxi Shan Exped. 61 (PE), Longxi Shan Exped. 1186 (PE); Jianning, Y.L. Gao 153 (FJSI), X.L. Hou & R.P. Zhang 1463 (AU); Nanping, G.S. He 1458 (FJSI), Y. Ling 2383 (PE), Wuyishan Exped. 801812 (FJIDC); Ningde, X.X. Su CSH15138 (CSH); Sanming, L.K. Ling 370 (PE); Shunchang, M.S. Li 5661 (FJSI); Taining, L.K. Ling 743 (AU, PE); Wuping, J.G. Lin & Z.Y. Lin 3698 (FJIDC); Wuyishan, G.S. He 2862 (FJSI), M.C. Wang 3252 (PE), M.C. Wang & B.X. Huang 3252 (NAS, PE), Wuyishan Exp. 2223 (FJSI), X. Zhong & Q.F. Zhang ZX03745 (CSH); Xiamen, Y. Ling 4880 (AU, PE); Zhenghe, Anonymous 4400 (FJIDC); Precise locality unknown, J. He 2156 (PE), J. He 2157 (PE). Gansu: Kangxian, C.Y. Chang 16707 (WUK); Tanchang, W.Y. Hsia 6763 (PE); Tianshui, K.M. Liou 10520 (PE); Wenxian, Baishuijiang Exped. 149 (PE), Baishuijiang Exped. 465 (PE), Baishuijiang Exped. 3283 (PE), Baishuijiang Exped. 3380 (PE), Baishuijiang Exped. 3419 (PE), Baishuijiang Exped.3425 (PE), Baishuijiang Exped.4019 (PE), Baishuijiang Exped.4259 (PE), Baishuijiang Exped. 4374 (PE), Baishuijiang Exped. 4431 (PE), Baishuijiang Exped. 4977 (PE), Baishuijiang Exped. 5163 (PE), Baishuijiang Exped. 5172 (PE), Baishuijiang Exped. 5354 (PE), Baishuijiang Exped. 5449 (PE), D.E. Boufford et al. 37593 (PE), L.Q. Li et al. 139 (PE), C.L. Tang 1683 (IBK, KUN, WUK), T.P. Wang 19228 (HNWP, WUK); Zhouqu, Bailongjiang Exped. 180 (PE), Bailongjiang Exped. 560 (PE). Guangdong: Lechang, S.H. Chun 3103 (IBK); Lianping, 236 Exped. 1220 (PE); Lianshan, P.C. Tam 58270 (IBK, KUN, PE); Ruyuan, S.P. Ko 52572 (KUN, NAS, PE); Xinyi, C. Wang 32209 (IBK, PE, SN); Precise locality unknown, S.S. Sin 9249 (PE). Guangxi: Guilin, L.Q. Chen 94345 (PE), Z.Z.Chen51042 (KUN,WUK), Guanyang Exped.450327160409002LY (IBK), Guanyang Exped.450327160718037LY (IBK), C.M. Hu & Q.H. Li 1496 (LBG), C.F. Liang 31187 (IBK), Lingui Exped. 450322170408002L (GXMG); Huanjiang, Beijing Exped. 893003 (PE), Beijing Exped. 895246 (PE), Huanjiang Exped. 451226130317015LY (IBK), Y.H. Liang 4‒3‒428 (GXMI); Jinxiu, Dayao Shan Exped. 10311 (IBK), Dayao Shan Exped. 12945 (IBK), D. Fang 64814 (GXMI), F.N. Wei 611 (IBK), R.B. Wei 14600 (GXMG); Lingchuan, Lingchuan Exped. 450323130312059LY (IBK); Lingyun, A.N. Steward & H.C. Cheo 175 (N); Longsheng, Z.S. Chung 91086 (IBK), Z.S. Chung 91108 (IBK), Guangfu Forest Gathering Team 12 (PE), Guangfu Forest Gathering Team 744 (IBK), B. Liu 542 (PE), Longsheng Exped. 450328140321016LY (GXMG); Luocheng, Beijing Exped. 895590 (PE), Luocheng Exped. 4‒1‒379 (GXMI); Nanning, L.Wu et al. D1808 (IBK); Quanzhou, Z.S. Chung 82065 (IBK), D.A. Huang 60636 (IBK), Q. Li 316 (PE), Quanzhou Exped. 450324130320043LY (GXMG, IBK); Rongshui, Beijing Exped. 892088 (PE), Beijing Exped. 892179 (PE), Beijing Exped. 892556 (PE), D. Fang & D.H. Qin 63934 (GXMI), Z.Y. Li et al. 891620 (HAST), Z.Y. Li et al. 892179 (HAST), R.S. Yang & Y.P. Tao 53‒59 (GXMI), Yuanbao Shan Exped. Y1028 (IBK); Tianlin, Tianlin Exped. 451029130419078 (GXMI); Xing’an, C.Z. Gao, H.Q. Wen & W.C. Yu 42312 (GXMI), F. Liu 213 (PE), G.Z. Li & B. Lu 10375 (IBK), G.Z. Li & B. Lu 12229 (IBK); Ziyuan, Ziyuan Exped. 450329160330017LY (IBK). Guizhou: Daozhen, X.M. Wang & P. Zhao 870434 (HGAS), X.M. Wang & P. Zhao 870508 (HGAS); Duyun, Guizhou Exped. 105 (HGAS), Y. Tsiang 5771 (NAS, PE), J. You DPS20115134 (QNUN), X.M. Zeng DPS20115255 (QNUN); Jiangkou, P.C. Tsoong 1819 (KUN), Y.L. Tu 93‒563 (GNUG), Z.Y. Zhang et al. 400149 (PE), Z.Y. Zhang et al. 400242 (HGAS, PE), Z.Y. Zhang et al. 400782 (PE), Z.Y. Zhang et al. 402072 (PE); Kaili, Qiannan Exped. 1140 (NAS), Qiannan Exped. 2029 (NAS, PE); Leishan, Q.H. Chen 3034 (HGAS), Q.H. Chen 3078 (HGAS), Q.H. Chen 3463 (HGAS), Q.H. Chen 3519 (HGAS), Q.H. Chen 74‒419 (HGAS), Qiannan Exped. 830 (PE), C.P. Tsien et al. 50224 (PE), D.W. Xie 3461 (HGAS); Libo, X.M. Wang 185 (HGAS); Liping, Q.H. Chen 826 (HGAS), Q.H. Chen 849 (HGAS, FJSI); Majiang, M.T. An 342 (GFS); Rongjiang, Qiannan Exped. 3281 (PE); Sandu, J.Z. Zhang QNSY 20041179 (QNUN); Shiqian, Q.H. Chen 94112 (HGAS), J. Huang 522224160402003LY (GZTM), Y. Tsiang 4165B (NAS); Songtao, Wuling Shan Exped. 88093 (PE); Suiyang, M.T. An et al. SY-1‒058 (GZAC), M.T. An et al. SY-20150230 (GZAC), X. Chen K0802240 (ZY), X.J. Chen 91111 (ZY), F.H. Li 66 (ZY), B. Liu 10 (ZY), B. Liu 2002 (ZY), T.T. Tan kks03028 (ZY), P.C. Tsoong 416 (PE); Tongzi, X.M. Wang & P. Zhao 870015 (HGAS); Yinjiang, M.T. An et al. YJ-2014‒0094 (GZAC), Qianbei Exped. 793 (HGAS, PE), Z.Y. Zhang et al. 401008 (HGAS, PE), Z.Y. Zhang et al. 401293 (PE), Z.Y. Zhang et al. 401838 (HGAS, PE); Zheng’an, H. Liu 520324160405014LY (GZTM); Zunyi, S.W. Deng 90146 (IBK), Fourth National Chin. Med. Exped. 520325160531627LY (GZTM), Qianbei Exped. 138 (HGAS, IBK), Qianbei Exped. 257 (HGAS), F.Y. Peng kks101189 (ZY), P.C. Tsoong 394 (PE). Hubei: Badong, H.M. Li & Y.S. Zhang 1083 (NAS); Hefeng, H.M. Li & Y.S. Zhang 1096 (NAS), H.J. Li 5686 (PE, WUK), F.S. Peng 561 (PE), Y.M. Wang 5709 (PE); Jianshi, L.Y. Dai & C.H. Qian 150 (NAS, PE), L.Y. Dai & C.H. Qian 1299 (NAS, PE); Laifeng, H.J. Li 7465 (IBK, PE, WUK); Shennongjia, Shennongjia Exped. 21077 (PE), Sino-Amer. W Hubei Plant Exped. 506 (KUN, NAS, PE), D.G. Zhang 5740 (JIU), D.G. Zhang lj0125190 (JIU); Wufeng, D.G. Zhang et al. 440 (HHE); Xingshan, H.M. Li & Y.S. Zhang 1089 (NAS), Y. Liu 422 (NAS, PE), M.X. Nie & Q.H. Li 422 (LBG, PE); Xuan’en, H.J. Li 887 (PE, WUK), H.J. Li 2707 (WUK), H.J. Li 3902 (PE), H.M. Li, Y.M. Yi & Y.S. Zhang 1077 (NAS), Y.M. Wang 4487 (PE); Yichang, H.B. Chen s.n. (FJSI), P.X. Dong 23 (PE), H.M. Li & Y.S. Zhang 1093 (NAS), R. Long & B.J. Zheng 90091 (PE); Zigui, J.Q. Zhang 240 (PE). Hunan: Baojing, X.J. Su & H.B. Liu 433125D00060505023 (JIU), D.G. Zhang 1144 (JIU), D.G. Zhang ly-128 (JIU); Chengbu, K.G. Li et al. CB20150514_0013 (JIU), S.D. Wu 19 (HUFD); Cili, W. Hunan Exped. 900 (PE); Guzhang, L. Su 4331261503031204 (JIU), D.G. Zhang 080712LLY002 (JIU), D.G. Zhang pws2694 (JIU), D.G. Zhang yd1144 (JIU), D.Z. Zhang YH060513915 (JIU), D.Z. Zhang YH150515793 (JIU), D.Z. Zhang YH990516916 (JIU), D.Z. Zhang zdg00015 (JIU); Fenghuang, Z.Q. Zheng & S.Q. Yang 120 (NAS); Hengyang, S.H. Chun 3257 (IBK), Y.Q. Kuang 691 (PE), J.L. Xia 201714 (NYA), C.S. Yok & C.M. Chen 1779 (NAS), C. Zhao 201006 (NYA); Hongjiang, C.T. Lee 248 (PE), C.T. Lee 1686 (PE), C.T. Lee 1758 (PE), C.T. Lee 2039 (PE), C.T. Lee 2589 (PE); Huaihua, P.C. Tam 60552 (IBK); Liuyang, Y.D. Liu 25609 (HNNU); Longhui, L.D. Duan 2115 (PE); Longshan, L.H. Liou 1641 (PE); Ruanling, Wuling Exped. 43 (PE); Sangzhi, Beijing Exped. 2002 (PE), Z.C. Gu 420027 (JIU), B.K. Lee & S.B. Wan 750023 (PE), C.M. Tan et al. 11 (JJF); Shimen, Hupingshan Exped. 796 (PE); Suining, J.J. Zhou & D. Zhou 143 (CSFI); Xinning, Anonymous 63443 (IBK), L.B. Luo 471 (PE), L.B. Luo 703 (PE), Y.B. Luo 204 (PE), Y.B. Luo 2033 (PE), Y.B. Luo 2048 (PE), Y.B. Luo 2049 (PE), Y.B. Luo 2162 (PE), Y.B. Luo 2339 (PE), Y.B. Luo 2449 (PE), Z.C. Luo 1330 (PE), Ziyunshan Exped. 223 (PE), Ziyunshan Exped. 1142 (PE), Ziyunshan Exped. 1157 (PE); Yongshun, Beijing Exped. 175 (PE), S.Q. Chun 729 (IBK), Hunan Exped. 292 (PE), K.D. Lei 4331271503051513 (JIU), D.G. Zhang zdg10239 (JIU), D.G. Zhang ZZ140305363 (JIU), Y. Zhang 72046 (JIU); Zhangjiajie, 83‒3 630275 (JIU), Z.C. Gu 82108 (JIU), H. Zhou & J.L. Luo 15030713 (CSFI), H. Zhou & J.L. Luo 15030717 (CSFI), H. Zhou & J.L. Luo 15030923 (CSFI), H. Zhou & D.S. Zhou 15040905 (CSFI); Zhuzhou, W.B. Liao, W.Y. Zhao & W.S. Yang LXP13‒5468 (SYS). Jiangxi: Anfu, Jiangxi Exped. 297 (PE), Y.G. Xiong 8116 (LBG), C.B. Yang & K. Yao 1046 (PE), C.S. Yok et al. 2740 (PE); Anyuan, Q.M. Hu & Q.H. Li 2005 (LBG); Chongwen, L. Li 20161104055 (GNNU); Guixi, C.M. Tan et al. 8757 (JJF); Ji’an, 236 Exped. 454 (PE); Jinggangshan, Anonymous (70)179 (PE), Q. Fan et al. JGS 4117 (SYS), S.S. Lai 84 (LBG), S.S. Lai, R.J. Yang & D.F. Huang 4406 (IBK, KUN, LBG), W.B. Liao, W.Y. Zhao & B.H. Wu LXP13‒15138 (SYS), C.S. Yok et al. 5031 (NAS, PE); Jiujiang, T.L. Bullock 180 ( US); Lichuan, X.X. Yang et al. 650008 (WUK); Lushan, Alpine Plant Team I‒298 (JSPC), K.C. Kuan 77054 (PE), S.S. Lai & S.C. Zhang 43 (LBG), H.M. Li, Y.S. Zhang & Y. Xu 1109 (NAS), H. Migo s.n. (NAS), M.X. Nie 92009 (HAST, PE), M.X. Nie & S.L. Chen 7110 (LBG, PE), C.M. Tan et al. 1507795 (JJF), F.Y. Wang wfy00203 (CSH), M.C. Wang & H. Zou 1764 (LBG), Y.G. Xiong 9921 (LBG, NAS, PE), G.H. Yi et al. 9684 (CCAU, SZG), G.F. Zhu 249 (LBG); Nanchang, M.X. Nie & S.S. Lai 2005 (PE, WUK); Nanfeng, M. X. Nie & S.S. Lai 2404 (LBG, PE), X.X. Yang et al. 650240 (WUK); Pingxiang, G.X. Chen & D.G. Zhang LXP‒064923 (JIU), H.J. Jing et al. LXP-GTY-260 (SYS), W.Y. Zhao et al. LXP-13‒09455 (SYS); Qianshan, Y.D. Chen & X.T. Ma 1134 (PE); Ruijin, Q.M. Hu 3563 (KUN, PE); Suichuan, J.J. Zhang, W.F. Cai & J.W. Pan LXP-13‒17262 (SYS); Wuning, C.M. Tan 9604081 (JJF, NAS, PE), C.M. Tan 97260 (JJF), C.M. Tan & J.S. Xiong 97241 (JJF), C.M. Tan & J.H. Zhang 97421 (JJF, SZG); Wuyuan, Q.H. Li & C. Chen 64 (PE); Xiushui, H.M. Li, Y.S. Zhang & Y. Xu 1100 (NAS); Xunwu, Q.M. Hu 1496 (PE), C.S. Yok et al. 1973 (PE); Yihuang, Q.H. Li & C. Chen 1600 (PE); Yongxiu, S.S. Lai 2211 (PE). Shaanxi: Hanzhong, X.X. Hou 377 (WUK), X.X. Hou 1212 (WUK); Lan’gao, Y.S. Chen et al. 2403 (WUK); Lueyang, K.T. Fu 5843 (IBK, KUN, PE, WUK); Pingli, Pingli Exped. 480 (WUK), G.Y. Xu 5635 (WUK); Yangxian, K.T. Fu 5279 (CDBI, IBK, PE, WUK). Sichuan: Baoxing, Baoxing Exped. 78‒0181 (SM); Bazhong, Bazhong Exped. 187 (SM); Cangxi, Anonymous 256 (SM), Anonymous 435 (SM), Z.S. Qin 2525 (CDBI); Chongzhou, Z.B. Feng, D.H. Zhu & X.J. Li 4337 (WCSBG), X.F. Gao, W.B. Ju & M. Li AZH00050 (CDBI), W.B. Ju & D.K. Chen AZH00653 (CDBI), W.B. Ju & D.K. Chen AZH00693 (CDBI), W.B. Ju, D.K. Chen & M. Li AZH00853 (CDBI), W.B. Ju, D.K. Chen & M. Li AZH00902 (CDBI), W.B. Ju, D.K. Chen & M. Li AZH00978 (CDBI), W.B. Ju, D.K. Chen & M. Li AZH01005 (CDBI), D.H. Zhu 4069 (HX); Dazhou, Dazhou Exped. 788 (SM); Dujiangyan, C.S. Chang et al. S10280 (PE), Z.B. Feng & H.M. Shao 20070003 (WCSBG), D.Z. Fu 87‒1473 (WCSBG), Y.Y. Geng et al. 981454 (PE, WCSBG), C. Zhang & Z.B. Feng 69 (WCSBG), C. Zhang 20060158 (HX), Z.L. Zhao 87‒118 (HX), Z.L. Zhao 87‒350 (HX), D.H. Zhu 3444 (WCSBG); Ebian, C.W. Yao 4362 (NAS), T.T. Yu 606 (PE), T.T. Yu 697 (PE); Emei Shan, H.M. Li & Y.S. Zhang 1157 (NAS), Sichuan Exped. 485 (CDBI, PEY), Sichuan Univ. Biol. Depart. Exped. 49440 (WUK), Q.S. Zhao & Z.M. Tan 116270 (CDBI); Ganluo, Anonymous 593 (SM); Gulin, Gulin Exped. 385 (SM), Gulin Exped. 667 (SM), Gulin Exped. 771 (SM); Hongya, W.K. Bao et al. 2494 (CDBI), H.M. Li & Y.S. Zhang 1160 (NAS), C.W. Yao 2398 (PE), C.W. Yao 3938 (PE); Jian’ge, Anonymous 67 (SM); Jianyang, Anonymous 2511 (SM); Leibo, Anonymous 28 (PE, SM), M.Y. Fang 117838 (CDBI), M.Y. He & Q.S. Zhao 116731 (CDBI), M.Y. He & Q.S. Zhao 116947 (CDBI), M.Y. He & Q.S. Zhao 117002 (CDBI, PE), C.T. Kuan 8693 (PE), Leibo Exped. 305 (SM), Leibo Exped. 312 (SM), Leibo Exped. 399 (SM), Sichuan Econ. Plant Exped. 730 (SM), Q.S. Zhao et al. 117617 (CDBI), Q.S. Zhao et al. 117645 (CDBI), K.P. Yin & Q.H. Chen 27 (CDBI), K.P. Yin & Q.H. Chen 46 (CDBI), K.P. Yin & Q.H. Chen 147 (CDBI), Q.S. Zhao & Z.M. Tan 116195 (CDBI); Mabian, Mabian Exped. 617 (SM), F.T. Wang 22952 (PE); Mianning, D.E. Boufford et al. 32966 (KUN); Mianyang, D.H. Zhu 3677 (WCSBG, WUK); Nanjiang, Anonymous 97 (SM), Y. He & H. Tian GWS201504057 (BNU), Y. He & H. Tian GWS201504093 (BNU), Y. He & H. Tian GWS201504098 (BNU); Pingwu, D.Z. Fu et al. 1488 (PE), D.Z. Fu et al. 1544 (PE), H.L. Tsiang 10232 (PE), H.L. Tsiang 10858 (PE), T. Wan et al. 1360 (PE), H.M. Yang et al. 1449 (PE), L. Zhang et al. 264 (PE), L. Zhang et al. 305 (PE), S.R. Zhang et al. 10 (PE), S.R. Zhang et al. 2132 (PE); Qingchuan, Anonymous 2194 (SM), Anonymous 2781 (SM); Qionglai, Anonymous 311 (SM), H.S. Kung 41881 (WUK); Shimian, C.C. Hsieh 40283 (PE); Tianquan, K.L. Chu 2299 (NAS, PE), D.Y. Peng 47869 (PE), Sichuan Econ. Plant Exped. 453 (PE); Tongjiang, Tongjiang Exped. 413 (SM); Wangcang, Bashan Exped. 4944 (PE); Wenchuan, Anonymous 683 (SM); Xuyong, Xuyong Exped. 354 (SM), Xuyong Exped. 610 (SM), Z.C. Zhong 1124 (SM). Taiwan: Chiayi, Ou & Kao 9182 (TAIF); Ilan, C.H. Lin 110 (HAST), C.H. Lin et al. 365 (HAST), C.H. Lin 566 (HAST), C.H. Lin & G.F. Zhong 218 (HAST), S. Sasaki s.n. (TAIF); Nantou, Y. Kudo & K. Mori 87 (TAIF), K. Sun et al. 11548 (KUN); Pingtung, J.C. Wang & K.C. Yang 9003 (HAST); Taichung, C.T. Huang 424 (HAST), C.I. Huang et al. 462 (HAST); Taoyuan, T.Y. Liu 1364 (HAST), C.I. Peng et al. 14635 (HAST), C.I. Peng et al. 22850 (HAST), K.Y. Wang 1137 (HAST). Yunnan: Wenshan, K.M. Feng 22234 (NAS), Y.M. Shui 383 (KUN), H.T. Tsai 51742 (NAS); Zhaotong, H.J. Dong, Z.H. Wang & Z.J. Yin WM-0191 (KUN, PE). Zhejiang: Chun’an, Zhejiang Plant Resour. Exped. 27702 (NAS); Hangzhou, P.L. Chiu 466 (HZ), M.B. Deng et al. 4607 (HZ, NAS, PE, WUK), Y.Y. Ho 21439 (HZ); Jiangshan, L.C. Jin & J.K. You 3570 (ZM); Lishui, C.F. Song, Y.N. Xiong & L. Zhao LDS2781 (NAS), C.F. Song, Y.N. Xiong & L. Zhao LDS2871 (NAS), C.F. Song, Y.N. Xiong & L. Zhao LDS2904 (NAS); Pan’an, F.G. Zhang DPS955 (ZM); Suichang, Chang & Li 5164 (ZM), R.H. Shan 6267 (NAS), Zhejiang Plant Resour. Exped. 25663 (HZ, ZM).

VIETNAM. Lào Cai: Sa Pa, P.A. Pételot 3703 (L).

Lectotypification:— Sanicula petagnioides was described on the basis of three collections, namely T. Kawakami & U. Mori 1988, T. Kawakami & U. Mori 2026, and G. Nakahara s.n. We have been able to trace three specimens of later two collections at TAIF (Fig. 3) and TI. All these specimens match well the protologue and are syntypes as Hayata (1908) did not designated a type for S. petagnioides . According to ICN Art. 9.11 ( Turland et al. 2018), here we designate T. Kawakami & U. Mori 2026 (TAIF18726; Fig. 3A), which is well preserved completely, as lectotype.

Notes:— In the protologue of Sanicula henryi, Wolff (1913) stated the collection A. Henry 10500 was preserved in B. However, it has most likely been destroyed during World War II. Another five duplicates are kept at E, K, MO, NY and US, respectively, and all well preserved and also agree with the original description. Pimenov (2017) designated K sheet as its lectotype, and the sheets at E, MO, NY and US are isolectotypes. According to ICN Art. 9.3 ( Turland et al. 2018), we accept the opinion of Pimenov (2017).