Atholus coelestis ( Marseul, 1857 )

Cruz, Ian Niel dela, Hartini, Sri, Dwibadra, Dhian & Ôhara, Masahiro, 2024, Contribution to the Knowledge of the Genus Atholus (Coleoptera: Histeridae: Histerinae: Histerini) from the Indonesian Archipelago, Species Diversity 29 (1), pp. 119-139 : 121-122

Atholus coelestis ( Marseul, 1857)

( Figs 1C , 11A )

Hister coelestis Marseul, 1857: 416 , pl. 10, fig. 59 (type locality: China) .

Hister femoralis Motschulsky, 1863: 449 [synonymized by Lewis (1885: 465)].

Atholus coelestis : Lewis 1906: 402 (listed); Lewis 1915: 55 ( Formosa); Mazur 1984: 212; Ôhara 1992: 173–176 (Nansei islands; redescription, male genitalia); Mazur 1997: 129; Ôhara 1999: 31–32; Mazur 2011: 104 (Oriental region, Tajikistan, Comores; catalogued); Mazur et al. 2015: 1454 ; Ho et al. 2023: 41–42 (Taiwan); dela Cruz and Ôhara 2023a: 1–26 ( Philippines); dela Cruz and Ôhara 2023b: 13–38 ( Japan).

Hister (Atholus) coelestis : Bickhardt 1910: 53 (catalogued); Bickhardt 1917: 193 (catalogued); Desbordes 1917: 320 (keyed); Desbordes 1919: 399 ( Tonkin, Annam, Cochinchine); Desbordes 1921: 10 ( Inde); Kamiya and Takagi 1938: 31 (listed).

Atholus (Euatholus) coelestis : Hisamatsu and Kusui 1984: 17 (noted, key).

Atholus (Euatholus) coelestes [sic]: Hisamatsu 1985: 228, pl. 41, fig. 61 (noted, keyed, imaged).

Specimens examined. Seventy-one specimens of undetermined sex. Sumatra Island: Central Sumatra, Payakumbuh , Harau Valley , November 1993, Sarimudanas leg . (2 exs SEHU, 1 ex. MZB) . Java Island : West Java, Sukabumi, Jampang, Tgl ., 9712001, Bojong Pari , 21–22 December 1997, alt . 640 m, E . and E . Cholik leg. (3 exs MZB); W. Java, 18. 3000, 20.30, Localitij, W . Preanger , November 1941, Groenendael leg . (1 ex. MZB); W . Java, Bandoeng , 24 January 1940, 700m, J . Olthof leg . (1 ex. MZB); Java, Preanger, N .O.I., Bandoeng , 25 February 1936, alt . 750m, F . C . Drescher leg . (1 ex. MZB); Java, Preanger N .O.I., Bandoeng , 12 February and 8 March 1936, alt . 750 m, F . C . Drescher leg . (4 exs MZB); Preanger, Bandoeng , 17 May 1932, F . C . Drescher leg . (1 ex. MZB); Tjiatj . [= Tjilatjap ], August 1925, unknown collector (1 ex. MZB) . Borneo Island : Kalimantan Timur, Kab . Kutai Kartanegara , 01.03042°S, 116.56526°E, Ds . Karya Merdeka RT 09, Kec, Samboja , 18 November 2007, M . Ôhara leg . [KL-07-MO-028] (1 ex. SEHU) . Sulawesi Island: Zuld Celebes, Nanggala , Rantepao , October 1937, alt . 900 m, F. C. Drescher (11 exs MZB); Nanggala , November 1937, unknown collector (4 exs MZB); Kabupaten Maros [near Makassar], 05.1008°S, 119.4038°E, 40 m . a.s.l., Cagar Alam Bantimurung , 27 October 2000, M . Ôhara leg . [SU-00-MO-002] (1 ex. SEHU, 1 ex. MZB); Kabupaten Barru, 04.2527°S, 119.3646°E, 2 m . a.s.l., Desa Tung, Kecamatan Barru , 28 October 2000, M . Ôhara leg . [SU-00-MO-005] (3 exs SEHU, 2 exs MZB); Kabupaten Tana Toraja, Kec . Makale, Tondon , 02.5744°S, 119.5442°E, 750 m . a.s.l., Pasar Hewan Bolu, Desa Bolu, Kecamatan Tallunglipu , 30 October 2000, M . Ôhara leg . [SU-00-MO-012] (1 ex. SEHU); Provinsi Sulawesi Selatan, Parepare, Mallusetasi, 04.0538°S, 119.3657°E, 3 m . a.s.l., Desa Bojo, Kecamatan Mallusetasi , 31 October 2000, M . Ôhara leg . [SU-00-MO-016] (3 exs SEHU, 2 exs MZB); Kab . Bolaang Mongondow , Kec . Dumoga Barat, 00.0118°N, 123.5633°E, Desa Doloduo, Kosinggolan , 1 September 2006, M . Ôhara leg . [SU-06-MO-054] (1 ex. SEHU); Kab . Bolaang Mongondow , Dumoga Barat, 00.3947°N, 124.0636°E, Desa Bumbungon, Kec . Dumoga , 2 September 2006, M . Ôhara leg . [SU-06-MO-057] (2 exs SEHU, 2 exs MZB); Provinsi Sulawesi Utara, Kab . Bolaang Mongondow , 00.5913°N, 124.1401°E, Desa Nonapan, Kec . Poigar , 2 September 2006, M . Ôhara leg . [SU-06-MO-059] (1 ex. SEHU); Kab . Maros , Kec . Bantimurung , Ds . Bontebonte , 04.5541°S, 119.0702°E, 3 September 2006, M . Ôhara leg . [SU-06-MO-063] (1 ex. SEHU); Kab . Maros , Kec . Simbang, Desa Samangki , 05.0243°S, 119.4243°E, 3 September 2006, M . Ôhara leg . [SU-06-MO-064] (1 ex. SEHU); Kab . Maros , Kec . Bantimurung, Desa Bantimurung , 05.0143°S, 119.4035°E, 3 September 2006, M . Ôhara leg . [SU-06-MO-065] (1 ex. SEHU); Sulawesi, Provinsi Sulawesi Selatan, Kab . Maros , Kec . Cenrana, Desa Limapoccoe , 5 September 2006, G . Takaku leg . [SU-06-GT-069] (1 ex. SEHU, 1 ex. MZB); Provinsi Sulawesi Selatan, Kab . Maros , Kec . Cenrana , Ds . Lebbotengae , 5 September 2006, G . Takaku leg . [SU-06-GT-070] (1 ex. MZB); Sulawesi Selatan, Bantimurung , 27 December 1999, N . Ohbayashi leg . [IC-22-02] (2 exs SEHU, 1 ex. MZB) . Bali Island: Sangeh , 10 February 1997, M . Kasagi leg . (1 ex. SEHU, 1 ex. MZB); 22 March 1981, K . Sugino leg . (1 ex. SEHU, 1 ex. MZB) . Sumbawa Island: Provinsi Nusa Tenggara Barat, Kabupaten Sumbawa Besar , Ropang , 08.5044°S, 117.1722°E, Hutan Malaka Desa Malaka , 9 November 2000, M . Ôhara leg . [SB-00-MO-033] (2 exs SEHU, 1 ex. MZB); Kabupaten Bima, Belo, 08.3713°S, 118.4324°E, Desa Ngali , Kecamatan Belo , 12 November 2000, M . Ôhara leg . [SB-00-MO-041] (2 exs SEHU, 1 ex. MZB) . Sumba Island: Provinsi Nusa Tenggara Timur, Kabupaten Sumba Timur, Kota Waingapu , 09.4055°S, 120.1219°E, Kecamatan Kota Waingapu , Kambajawa , 30 January 2003, M . Ôhara leg . [SM-03-MO-020] (1 ex. SEHU) .

Diagnosis. Atholus coelestis is characterized by its third dorsal stria extending inwardly towards the apical end of the fourth and fifth striae. Sometimes, the apex of the fourth stria also extends slightly towards the fifth stria. Among all the Indonesian Atholus , A. coelestis is discriminated for its slight emargination on its anterior margin of mesoventrite.

Distribution. China, Indochina, Sri Lanka, Nepal, Tajikistan, Comoros, Japan, Taiwan, Philippines, Indonesia: Greater Sunda (Sumatra, Java, Borneo and Sulawesi Islands), Lesser Sunda (Bali, Sumbawa and Sumba Islands) ( Fig. 11A ).

Remark. The slight emargination of on the anterior margin of its mesoventrite is unique to A. coelestis among all the Oriental and East Palearctic Atholus .

Figures

Fig. 1. Some Indonesian Atholus, dorsal habitus. A, Atholus bifrons (Marseul, 1854), IC-22-in01; B, A. philippinensis (Marseul, 1854), IC- 22-in06; C, A. coelestis (Marseul, 1857), IC-22-in02; D, A. myrmidon (Marseul, 1861), IC-22-in05; E, A. famulus (Lewis, 1892), IC-22-in03; F, A. singalanus (Marseul, 1880), IC-22-in07; G, A. torquatus (Marseul, 1854), IC-22-in08. Scale bar: 1.00mm.

Fig. 11. Map showing the occurrence of some species of Atholus from the Indonesian archipelago and its extra-limital distribution in the Oriental Region. A, Atholus coelestis; B, A. philippinesis; C, A. torquatus; D, A. bifrons.