Atholus, Thomson, 1859

Cruz, Ian Niel dela, Hartini, Sri, Dwibadra, Dhian & Ôhara, Masahiro, 2024, Contribution to the Knowledge of the Genus Atholus (Coleoptera: Histeridae: Histerinae: Histerini) from the Indonesian Archipelago, Species Diversity 29 (1), pp. 119-139 : 120

Key to the Indonesian Species of the Genus Atholus (Modified from Desbordes 1917), Excluding A. mundulus

1. Lateral pronotal striae two............................................... A. tenuistriatus ( Lewis, 1889)

— Lateral pronotal stria one......................... 2

2. Apical end of third dorsal stria strongly bent inwards ......................... A. coelestis ( Marseul, 1857)

— Third dorsal stria of elytra not bent inwardly........ 3

3. Lateral pronotal stria complete, continued as anterior stria behind the head............................. 4

— Lateral pronotal stria interrupted in the anterolateral angle.......................................... 8

4. Elytral dorsal striae 1–3 entire, dorsal stria 4 present only on apical end............................... 5

— Elytral dorsal striae 1–4 complete.................. 6

5. External subhumeral stria of elytra long, slightly abbreviated on apical fourth............................................... A. philippinensis ( Marseul, 1854)

— External subhumeral stria of elytra short, reduced to an arc, present on basal end................................................. A. singalanus ( Marseul, 1880)

6. Pygidium strongly punctate................................................. A. bifrons ( Marseul, 1854)

— Pygidium smooth or finely punctate............... 7

7. External subhumeral stria of elytra very short or absent; prosternal lobe punctate............................................. A. terraemotus ( Lewis, 1900)

— External subhumeral stria of elytra well-marked on medio-basal portion; prosternal lobe smooth............................... A. myrmidon ( Marseul, 1861)

8. Pronotal foveae behind anterolateral angle present............................... A. tetricus ( Lewis, 1902)

— Anterolateral area of pronotum without foveae...... 9

9. Propygidium with shallow excavation with coarse punctures inside, occupying middle and a little more than the posterior half..... A. gestroi ( Schmidt, 1897)

— Propygidium normal, with no conspicuous excavation on the surface.................................. 10

10. Apex of lateral pronotal stria arched behind the anterior angle...................................... 11

— Apex of lateral stria of pronotum not arched behind the anterior angle.................................. 12

11. Lateral stria of pronotum more shortened basally and less hamate on the apical end; pygidium smooth................................ A. pinnulae ( Lewis, 1900)

— Lateral stria of pronotum less shortened basally and more hamate on the apical end; punctures in pygidium becoming finer apically... A. torquatus ( Marseul, 1854)

12. Anterior margin of clypeus with strong indentation, the anterior edge before the eye deeply notched; pygidium with irregular and sparse punctures.................................... A. crenatifrons ( Lewis, 1899)

— Anterior margin of clypeus normal, feebly sinuous laterally; pygidium entirely smooth............................................. A. famulus ( Lewis, 1892)