Cossyphodes naukluftensis, Schawaller, Wolfgang, 2013

Schawaller, Wolfgang, 2013, Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue, Zootaxa 3721 (4), pp. 351-364 : 359

Cossyphodes mourgliai Ferrer, 1993 (in Ferrer & Collingwood 1993)

( Fig. 22 )

Cossyphodes naukluftensis sp. n. ( Fig. 10 )

Type material. Holotype (sex unknown): Namibia (labelled as S. W. Afr.), Naukluft Park, 24°16'S / 16°15'E, 26.X.1974, leg. S. Endrödy-Younga, TMSA.

Description. Body reddish brown without colour pattern. Body length 2.3 mm. Head semicircular, clypeus not separated, clypeal lines indistinct, straight; anterior and lateral margins somewhat bent upwards, frons with a pair of distinct, complete external keels and a pair of a weak internal keels, anterior of internal keel with tubercle; head surface with regular microgranulation; eyes narrow sickle-shaped, composed by two rows of dark ocelli, each row with about five ocelli; antennae 11-segmented with two large apical antennomeres forming club. Pronotum 1.5 times broader than median length, posterior corner rectangular, acute, anterior corner rounded; all margins unbordered; surface with same microgranulation and microsetation as on head, disc on each side with three complete longitudinal keels, additionally with distinct medial keel, this medial keel shortly interrupted in anterior part, for direction, distances of these keels, see Fig. 10 ; prosternal process elongate triangular. Elytra widest shortly before base, 1.4 times longer than broad; elytra with four primary and secondary keels, internal main keel only present near base of elytra, all main keels convergent from base towards tip, for direction, distances of these keels, see Fig. 10 ; surface with the same microgranulation and microsetation as on head and pronotum. Legs without peculiarities, tarsal formula 5-4-4. Aedeagus unknown, sex not examined (see introduction).

Diagnosis. C. naukluftensis sp. n. is similar to C. arnoldi Brauns, 1925 , known from Zimbabwe ( Fig. 9 ). Both share the narrow body shape, the head with a pair of distinct keels and with a tubercle before the internal keel, and the distinct pronotal keels with interrupted medial keel. Both species can be separated by a completely different pattern of main elytral keels ( Figs. 9–10 ). In C. naukluftensis sp. n. the elytral keels are bent outwards near base and convergent from base towards tip, in C. arnoldi the main keels are parallel in anterior half of elytra and convergent only in posterior half; in C. naukluftensis sp. n. the internal main keel is present only near base, in C. arnoldi this keel is complete.

Etymology. Named after the Naukluft Mountains, where the holotype was collected.

Figures

FIGURES 21 – 26. Dorsal view of African Cossyphodini. 21. Cossyphodes raffrayi Gestro, 1874, Tanzania, non-type HNHM. 22. Cossyphodes mourgliai Ferrer, 1993, Zambia, holotype ZSM. 23. Cossyphodes uviranus Basilewsky, 1951, Congo, nontype ZFMK. 24. Cossyphodes vandami Andreae, 1961, RSA / Pretoria, paratype MZLU. 25. Paramellops bewicki (Wollaston, 1861), RSA / Matjiesfontein, non-type TMSA. 26. Cossyphodites woodroofei (Péringuey, 1885), South Africa, non-type HNHM. Scale: 1 mm.

FIGURES 7 – 13. Dorsal view of African Cossyphodini. 7 – 8. Cossyphodes andreaei sp. n., RSA / Waterberg, paratype SMNS and aedeagus. 9. Cossyphodes arnoldi Brauns, 1925, Zimbabwe, lectotype TMSA. 10. Cossyphodes naukluftensis sp. n., Namibia / Naukluft, holotype TMSA. 11. Cossyphodes angolanus Basilewsky, 1952, holotype MRUC. 12. Cossyphodes wasmanni Reichensperger, 1915 (Cossyphodes brincki Andreae, 1961 syn. n., RSA / Hluhluwe, holotype MZLU). 13. Cossyphodes wasmanni Reichensperger, 1915, Zimbabwe, lectotype TMSA. Scale: 1 mm, for aedeagus: 0.5 mm.