Triphysa nervosa nervosa Motschoulsky, 1866

Dubatolov, Vladimir V., Korb, Stanislav K. & Yakovlev, Roman V., 2023, A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae), Ecologica Montenegrina 61, pp. 88-123 : 102-104

Triphysa nervosa nervosa Motschoulsky, 1866

Figs. 5i–p , 6a–b , 19 , 32, 34 .

= Triphysa phryne yonsaensis Im, 1988 ; Bull. Acad. Sci. P. R. Korea 1988 (3): 48-49.

Type locality: Democratic People's Republic of Korea ( North Korea), " North Hamgyong, Yonsa County ,

Sampori" (original spelling: “ ăŜ북ΞάήΞΘŀö ”).

Diagnosis. Male wings upperside with almost no white border. Wings underside with pale veins, but without postdiscal spots. Light strokes are located between the veins on hindwing underside. Few males from a series collected at the left bank of Budyumkan river in Chita Prov. have light areas in places where the eyespots should be present.

Material examined. 1 female, Malaya Sos’va Reserve, Northern Ural, estuary of Potlokh [Potlokh- Yugan] river, 14.VI 1989 ; 3 males, 1 female, Russia, Chita reg., near Naminga vill., 4-VII-2002, leg. V. Ivonin ( RYB) ; 2 males, Russia, Chita reg., Uryupino ( ISEAN) ; 1 male, 1 female, Sibiria or. (Irkutsk), 16/ 29- V-1910, O. Hesse ( ZMHU) ; 1 male, 1 female, Chita Prov., Argun’ river, Uryupino , 26-V-2001, V.V. Dubatolov, S. Gordeev, T. Vlasova ( ISEAN) ; 1 male, 1 female, Chita Prov., left bank of Budyumkan river, 5–7 km up from source, a slope with Quercus mongolica, 26-V-2001, V.V. Dubatolov, S. Gordeev ( ISEAN) ; 3 males, 1 female, 29-V-2001, V.V. Dubatolov, S. Gordeev ( ISEAN) ; 1 male, 1–2 km N from Budyumkan river source, swampy stream valley on a plateau, 31-V-2001, V.V. Dubatolov, S. Gordeev ( ISEAN) ; 6 males, 5 females, Russia, Chita reg., left bank of Gazimur river, near Kundara vill., 25-V-2001 ( ISEAN) ; 1 male, 1 female, SW Yakutia, Dakhabylskaya Steppe, river Delinde (in Chona river ), Tkachenko, 8.VII.[19]26 ( ZISP) ; 4 males, Yakutia, Oimyakon distr., Ust’-Nera settl., 16-VI-2010, leg. A. Samus ( ASV) ; 1 male, 180 km ENE Khandyga, Verkhnaya Khandyga upper course, 232 km of the road Khandyga – Magadan, Kuberlyakh stream valley, 3-VII-1985, V.V. Dubatolov ( ISEAN) ; 1 female, 180 km ENE Khandyga, Verkhnaya Khandyga upper course, 232 – [23] 5 km of the road Khandyga – Magadan, swamp, 28-VI-1985, V.V. Dubatolov ( ISEAN) ; 1 male, Yakutia, 234 km of the road Khandyga-Magadan, Vostochnaya Khandyga river upper course, 24-VI-[19]85, L.Popova ( ISEAN) ; 1 male, Yakutia, 300 km ENE Khandyga, Suntar river upper course, Indigirka river basin, 376 km [of the road Khandyga - Magadan], valley, right bank, meadow, 19-VII-1985, V.V. Dubatolov ( ISEAN) ; 1 male, Yakutia, 115 km NE Ust-Nera, Ytabyt-Yuryakh river, 10-30 km upper its source (left tributary of Indigirka river ), 5-VII-1987, I.V. Dorofeev ( ISEAN) ; 3 males, 1 female, Yakutia, Khandyga , 12-VI-1985, V.V. Dubatolov ( ISEAN) ; 1 male, 1 female, Yakutia , 232 km road Khandyga-Magadan ( ISEAN) ; 4 males, Yakutia, 270 km ENE Khandyga river, Kyubyume stream, Kyubyume settlement, Indigirka river basin, 336,5 km [of the road Khandyga - Magadan], 18-VII-1985, V.V.Dubatolov ( ISEAN) ; 3 males, 1 female, Yakutia, 364 km road Khandyga-Magadan, Suntar river , 20- VII-[19]85, L. Popova ( ISEAN) ; 1 male, Yakutia, 115 km NE Ust’ - Nera, Ytabyt-Yuryakh river ( ISEAN) ; 4 males, N. Yakutia, Ege-Khaja vill., 20-VI-89 ( ENH) ; 1 male, 1 female, Amur ( ZMHU) ; 1 male, 1 female, Amur reg., Blagovestshensk , 17-V-1995, leg. A. Streltsov ( RYB) ; 1 male, Amur reg., Svobodnyi distr., Malaya Sazanka vill., 25-V-1999, leg. A. Streltsov ( RYB) ; 1 male, Sokhondinsky Nature reserve, lower than Verkhny Bukukun, a road in Larix forest , 21-VI-1991, V.K. Zinchenko ( ISEAN) ; 2 males, Sokhondinsky Nature reserve, Bukukun river valley, kordon Ernisty , 4-5-VI-1991 ( ISEAN) ; 1 male, upper course of Chita river , 16-VI-[19]39, Cherepanov ( ISEAN) ; 1 male, Amur reg., 80 km N Urusha vill., 18-VI-1999, leg. A. Streltsov ( RYB) ; 2 males, Amur reg., Urusha distr., 7 km NE from Khalan Mt., Khaimkan river ( ISEAN) ; 1 male, Amur reg., 40 km NW Urusha ( ISEAN) ; 1 male, NW Amur reg., Urushinsky Mts., 7–8 km NE Khalan Mt., Khaimkan stream valley, 18-VI-1999, V.V. Dubatolov ( ISEAN) ; 6 males, 1 female, same locality, 21- VI-1999, V.V. Dubatolov ( ISEAN) ; 1 female, Amur reg., 40-45 km NW Urusha, Amutkachi village , 18-VI-1999, V.V. Dubatolov ( ISEAN) ; 1 female, Amur reg., Urusha environs, 18-VI-1999, V.V. Dubatolov ( ISEAN) ; 1 female, Zeya Nature Reserve , ”Goltzy” kordon environs, tundra, 54"07'N, 126" 56'E, 24-VI-2011 ( ISEAN) ; 1 female, Amur reg., Malaya Sazanka vill., 10-VI-[19]96, A.N. Streltzov ( ISEAN) ; 1 male, Blagovestshensk , environs, 25-V-[19]94, A.N. Streltzov ( ISEAN) ; 1 male, same locality, 21-VI-1996, A.N.Streltzov ( ISEAN) ; 1 female, western environs of Blagovestshensk, Verkhneblagovestshenskoe , agrobiological camp environs, 27-V-1999, V.V. Dubatolov ( ISEAN) ; 1 male, 1 female, Amur reg., Kuvykta station , 19-VI-1996, A. Streltzov ( ISEAN) ; 2 males, Amur ( MHUB) ; 1 male, Schilka ( MHUB) ; 1 male, 2 females, Pokrofka ( MHUB) ; 10 males, Khabarovsk reg., Dusse-Alin’ Mts., Pravaya Bureya riv. valley, kordon “Medvezh’e”, 16-VI-2012, E. Novomodnyi leg. ( ENH) ; 1 male, Khabarovsk Prov., Pivan’ environs, 21-VI-[19]77, V.A.Mutin ( ISEAN) ; 2 males, 1 female, Khabarovsk Prov., Khekhzyr , 10-VI-1980, [Chulkov] ( ISEAN) ; 1 female, Khabarovsk reg., Chegdomyn vill., park, 5-VI-2003, E. Novomodnyi leg. ( ENH) ; 8 males, 4 females, Khabarovsk reg., Bol’shoj Khekhzir Mts., Chirki , 22-VI-1987, E. Novomodnyi leg. ( ENH) ; 2 males, Khabarovsk reg., Berezovyi vill., BAM, 25–27-VI-1973, leg. Dolgikh ( ENH) ; 2 males, Khabarovsk reg., Evoron lake , 16–19-VI-1973, leg. Dolgikh ( ENH) ; 2 males, Khabarovsk reg., Sidima vill., 13-VI-1981 ( ENH) ; 1 male, 1 female, Middle Sikhote-Alin’ Mts., Zolotoi vill., Nemiz riv., 12-VI-1981 ( ENH) ; 2 males, 1 female, Malyi Khingan Mts., Kuldur , 1965, leg. Chulkov ( ENH) ; 1 male, 2 females, Amur, Zimmermanovka vill., 25-VI-1987, E. Novomodnyi leg. ( ENH) ; 1 female, Primorye reg., Chuguevka , 10-V-1993, leg. Yu. Gluschenko ( RYB) ; 1 male, Magadan reg., Omolon Sovkhoz, Kegali river , 3-VII- [19]68, Gomoyunova ( ISEAN) ; 1 male, Magadan Prov., 23-VIII-[19]68, Levina ( ISEAN) ; 2 males, Magadan Prov., Kegali , 30-VI-[19]69, V. Nikolaev ( ISEAN) ; 1 male, Magadan Prov., Palatka settlement env., 5-VII-1997, V.V. Papekha ( ISEAN) ; 2 males, Magadan reg., Aborigen station, Mai-Urusta , 26-VI-1997 ( RYB) ; 2 males, 1 female, Magadan reg., Omsukchan distr., Kupka, 119 km road to Omsukchan , 700 m, 61°59’43”N, 153°56’07”E, 28-VI-2006, leg. Cernila ( MCK) ; 1 male, 1 female, Chukotka, 10 km from Markovo, Anadyr’ valley , 28-VI-[19]67, Mirzaeva ( ISEAN) ; 1 female, 14.VII.1894, Mongolia Terilgje Kaschkaroff, coll. G.D. Nikolai Mikhailovich ( ZISP) ; 1 female, Nertschinsk , 25.VI ( ZISP) ; 1 female, Nertschinsk , Fabri coll., 31.VI.[1]907 ; 1 male, 1.VIII.[1]907; 1 female, 2.VIII.[1]907; 1 male, 15.VII.[1]908; 1 female, 31.VII [1]908 ( ZISP) ; 1 female, Nertschinsk, Sukhoi Log , 5.VI.1906 ; 1 female, 21.VI.1907; 1 female, 1.VII.[1]909; 1 female, 10.VI.[1]910; 1 male, 14.VI.[1]910; 1 male, 21.VI.[1]910; 2 males, 24.VI [1]910, Fabri coll.; 1 male, 14.VI.[1]910 ( ZISP) ; 1 male, Sretensk, Zabaik. , 10.VI.1930, Kapustin ( ZISP) ; 2 males, 4 females, Pochrofka , coll. Dieckmann, Graeser legit. ( ZISP) ; 1 male, Albasin, Puzilo , VII/VIII coll. Ersch. [off] ( ZISP) ; 2 males, Bol’shoi Never Amursk. Prov., Zakharzhevsky , 15.VI.[1]927 ( ZISP) ; 3 males, 1 female, Tukuringra-Gebge. Amur-Geb. Koshantschikow 22.V.[1]914 ( ZISP) ; 5 males, Seja Amur-Geb. Koshantschikow 20.V.1914 ; 2 males, 1 female, 22.V.[1]914; 1 male, 1 female, 23.V.1914 ( ZISP) ; 1 male, Amur, st. Kozakevicheva [Kazakevichevo, Ussuri valley], 1897, Bykov ( ZISP) ; 3 males, Koslofska [Kozlovka, near Bikin, Ussuri valley], Bykow ( ZISP) ; 1 male, Indigirka Valley, near Zashiversk, 30.VI- [19]20, Indig. [irka] Exp. [edition] Ak. [ademy] S.[cience] ( ZISP) ; 1 female, Sr.Kol. [ Srednekolymsk ], 29.VI.[1]905 ( ZISP) ; 1 male, 1 female, W. Chukotka, Bilibino , 5.VII.[19]90, V. Sinyaev ( ZISP) .

Distribution. Russia: Chita reg. (Sokhondo Mts., Udokan Range, Argun river lower course) ( Dubatolov & Gordeev 2002), Amur Prov., Khabarovsk Prov. ( Kurentsov 1970; Mutin 1992; Koshkin et al. 2007), Ussuri Prov. ( Martynenko 2000), Western, Southern and North-Eastern Yakutia ( Herz 1903a, 1903b; Kajmuk et al. 2005), Magadan reg. ( Takahashi & Oshima 2005), W. Chukotka (Markovo, Bilibino); North Korea; and in NE China ( Nikitin 1945; Tuzov 1995). The isolated population of Triphysa nervosa recorded by Gorbunov (1992) from the Malaya Sos’va nature reserve (NW Siberia, Khanty- Mansi Autonome District, Russia) and from the environs of Taezhnyi village (on the border between Khanty-Mansi Autonome District and Sverdlovsk reg., Russia). Another isolated population is known from the Sokhondo mountains in south-western part of Chita Prov.; this population is surrounded by other subspecies occuring in mountain hollows.

Figures

Figure 5. Triphysa sp., imago: a – T. nervosa brinikhi, holotype (ISEAN), upperside, b – underside; c – T. nervosa brinikhi, paratype, female (ISEAN), upperside, d – underside; e – T. nervosa tscherskii, male, lectotype (ZISP), upperside, f – underside; g – T. nervosa tscherskii, female, Yakutsk, 29.05.1985, V. Dubatolov leg. (ISEAN), upperside, h – underside; i – T. nervosa nervosa, male, Russia, Chita reg., near Naminga vill., 4.07.2002, leg. V. Ivonin (RYB), upperside, j – underside; k – T. nervosa nervosa, male, Amur reg., Blagoveshensk, 17.05.1995, leg. A. Streltsov (RYB), upperside, l – underside; m – T. nervosa nervosa, female, Amur reg., Blagoveshensk, 17.05.1995, leg. A. Streltsov (RYB), upperside, n – underside; o – T. nervosa nervosa, male, N. Ural, Malaya Sos’va Reserve, 14.06.1989 (ISEAN), upperside, p – underside.

Figure 6. Distribution map of Triphysa species: a (above) – general map; b (below) – northern-eastern part of area of T. nervosa nervosa.

Figures 18–19. Habitats of Triphysa: 18 – T. nervosa tscherskii, near Yakutsk (photo by A. Burnasheva); 19 – T. nervosa nervosa, E. Yakutiya, near Ust’-Nera vill. (photo by A. Samus’).

Figures 31–34. Male genitalia (lateral projection) of Triphysa: 31 – T. nervosa brinikhi, holotype; 32 – T. nervosa nervosa, Amur reg., Blagoveshensk; 33 –T. nervosa tscherskii, Yakutsk; 34 – T. nervosa nervosa, Magadan reg.