Triphysa Zeller, 1850

Dubatolov, Vladimir V., Korb, Stanislav K. & Yakovlev, Roman V., 2023, A review of the genus Triphysa Zeller, 1858 (Lepidoptera, Satyridae), Ecologica Montenegrina 61, pp. 88-123 : 91

Genus Triphysa Zeller, 1850

Entomol. Zeitung, Stettin 11: 308.

Type species: Papilo tircis Stoll, 1782 ( Hemming 1967: 449).

Diagnosis of the genus

Medium size butterflies, forewing length 17.0–23.0 mm. Wings in males dark, brown or grey; wings in females white. Upperside with characteristic light suffusion along veins. Black spots with white centers present at least in forewing upperside in males; in underside these spots are always present in both wings. In the male genitalia tegumen is domed, uncus longer than tegumen, apex of uncus rounded. Valva wide, apex of valva pointed. Aedeagus longer than valva, with characteristic large spikes on its apical part. In the female genitalia the signum shorter than 1/4 of bursa copulatrix (in Coenonympha s.l. and Lyela it is about 1/2 of its length).

From all genera of Coenonymphini Triphysa differs by the following features:

• In female genitalia signum shorter than 1/4 of bursa copulatrix;

• In male genitalia aedeagus with spikes in its apex;

• In male genitalia valva is wide (width/length = 2/1);

• Tegumen domed;

• Gnathos 2 times or less shorter than uncus;

• Males and females have opposite wing coloration (males dark, brown or grey, females light, white);

• Upperside of wings with characteristic light suffusion along the veins.