Mycomya goral, Väisänen, Rauno, 2013

Väisänen, Rauno, 2013, New Mycomya species from the Himalayas (Diptera, Mycetophilidae): 3. Subgenera Cesamya and Mycomyopsis, Zootaxa 3737 (2), pp. 129-153 : 133-137

Mycomya goral sp. n.

Figs. 2 A–F

Material studied. Holotype. ♂. NEPAL, Ktmd. Godavari, 6000', 17.VII. 1967, Can Nepal Exp. (in CNC). Paratypes (only ♂♂). Same data, 1 ♀ (in CNC); same data, but Mal. tr., 21.VII. 1967, 1 ♂, 1 ♀ (in CNC); same data, but Mal. tr., 2.VIII. 1967, 2 ♂♂ (in CNC); same data, but 3.VIII. 1967, 2 ♂♂ (in CNC and MZH).

Description. ♂. Head. Palp yellow, basally yellowish to brownish, other mouthparts yellow, face yellow, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1st flagellomere yellow. Length of 1st flagellomere about 3.5x, 2nd flagellomere 2x, its width. Thorax. Pronotum yellow, with 2 long setae. Scutum yellowish to brownish with 3 indistinct brownish to brown longitudinal, posteriorly fused stripes, anterolateral corners of scutum yellow. Anepisternum yellow. Preepisternum yellow. Scutellum light brownish, with 4 long setae. Laterotergite light brownish. Mediotergite light brownish, bare. Wing. Length 3.2–3.5 mm. Wing hyaline. Sc ending in R1 near or distad of middle of small cell, Sc1 apically broken or entirely missing. Apical part of Sc bearing 6–13 small setae. Small cell 1– 2 x as long as wide. Cu fork distinctly distal to M fork. M ratios: 0.95–1.05, 1.38–1.54. Cu ratios: 0.84–0.95, 1.40–1.67. Small setae: M petiole: 0; M1: 0–5; M2: 0–1; Cu petiole: 0; Cu1: 0; Cu2: 0. Halter pale yellowish, apex yellowish to brownish. Legs. Coxae yellow, femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.74–0.75, bt2:t2 = 0.63–0.67, bt3:t3 = 0.56–0.60. Abdomen. Tergites brownish to brown, sternites slightly paler. Hypopygium. Figs. 2 A–F, yellowish. Tergal part with 1 wide median comb of short spines, and 2 narrow inner combs, both with about 5 spines. Dark inner spur of tergal part apically rounded ( Fig. 2 C). Tergal lateral appendage setose, slender, about 5x as long as wide, with 2 flattened, curved subapical setae ( Fig. 2 D). Sternal submedian filament long, slender, curved, extending far beyond apex of aedeagus. Gonostylus about 5x as long as wide, with 3 subapical teeth, 1 strong seta, and membranous lateral lobe more than ½ of length of main branch of gonostylus ( Fig. 2 E). Aedeagus with 2 lateral lobes, extending slightly beyond its apex ( Fig. 2 F). Female. Wing length 4.0 mm. Thorax yellow, scutum with brown medial stripe, yellowish submedial stripes and brownish sublateral stripes, anterolateral corners of scutum yellow, mediotergite and laterotergite light brownish. Abdomen light brownish. Leg ratios: bt1:t1 = 0.83, bt2:t2 = 0.67, bt3:t3 = 0.60. Terminalia yellow.

Discussion. Mycomya goral is a small species and closely related to M. niltava sp. n. and M. sachak sp. n., both of which have a similar wide tergal comb of small spines, a similar general shape of the tergal lateral appendage ( Figs. 2 D, 3E, 4D), and long curved sternal submedian filaments. Mycomya goral differs from M. sachak in having an isolated flattened spine in the middle part on the inner margin of the tergal lateral appendage ( Fig. 2 D) similar to that in M. niltava ( Fig. 3 E). See also discussion under M. niltava .

Etymology. The species name is a noun in apposition and refers to the Himalayan goral , Nemorhaedus goral (Hardwicke) , a bovid species found across the Himalayas.

Mycomya niltava sp. n. Figs. 3 A–G

Material studied. Holotype. ♂. NEPAL, Dobang Kharka, 2400 m 83°24'E. 28°36'N. Oct. 28 1971 A. Nakanishi (in KUC). Paratypes (only ♂♂). Same data as holotype, 2 ♀♀ (in KUC); same data, but 9.X. 1971, 4 ♂♂, 6 ♀♀ (in KUC); same data, but 19.X. 1971, 1 ♀ (in KUC); same data, but 20.X. 1971, 28 ♂♂, 79 ♀♀ (in KUC); same data, but 23.X. 1971, 1 ♂, 1 ♀ (in KUC); same data, but 30.X. 1971, 1 ♂ (in KUC); Lipshiba Kharka, 83°23'E, 28°35'N, 1900 m, 30.X. 1971, A. Nakanishi, 1 ♀ (in KUC).

Description. ♂. Head. Palp yellowish, other mouthparts and face brownish, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1st flagellomere yellow. Length of 1st flagellomere about 4x, 2nd flagellomere 2.5x, its width. Thorax. Pronotum, with 3 long setae. Scutum yellowish with 3 brownish longitudinal stripes. Anepisternum yellowish to brownish. Preepisternum brownish, dorsal part lighter. Scutellum brownish, with 4 long setae. Laterotergite brownish, dorsal part lighter. Mediotergite brownish, bare. Wing. Length 3.9–4.3 mm. Wing hyaline. Sc ending in R1 near or distad of middle of small cell, Sc1 missing or very short. Apical part of Sc bearing 4–15 small setae. Small cell 1–1.5x as long as wide. Cu fork distinctly distal to M fork. M ratios: 0.85– 1.04, 1.15–1.41. Cu ratios: 0.89–1.00, 1.47–1.80. Small setae: M petiole: 0; M1: 0; M2: 0–1; Cu petiole: 0; Cu1: 0; Cu2: 0. Halter pale yellowish, apex brownish. Legs. Coxae and femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.77–0.83, bt2:t2 = 0.67–0.71, bt3:t3 = 0.57–0.64. Abdomen. Tergites brownish to brown, sternites yellowish. Hypopygium. Figs. 3 A–G, yellowish. Tergal part with 1 wide outer comb of spines, and 2 inner combs, both with about 7 spines. Almost whole base of outer comb widely setose, only its narrow median part bare. Dark inner spur of tergal part apically pointed ( Fig. 3 D). Tergal lateral appendage setose, slender, curved, about 6x as long as wide, apically slightly wider, with 1 flattened seta on median side of middle part and some flattened, curved apical setae ( Fig. 3 E). Gonostylus very long and slender, at least 8x as long as wide, with 2–3 subapical teeth and small membranous lateral lobe of 1/3 of length of main branch of gonostylus ( Fig. 3 F). Sternal submedian filament long and curved, extending far beyond apex of aedeagus. Aedeagus with 2 lobes, both with curved, spur-like group of setae. Female. Wing length 4.0–5.0 mm. Thorax as in ♂. Abdomen brownish to brown, sternites lighter, posteriorly yellow. Leg ratios: bt1:t1 = 0.77–0.81, bt2:t2 = 0.69–0.73, bt3:t3 = 0.53–0.63. Terminalia yellow.

Discussion. Mycomya niltava has a conspicuous, isolated and flattened seta on the inner margin of its tergal lateral appendage ( Fig. 3 E) resembling that of M. goral sp. n., which is a smaller species having a short gonostylus with a large lateral lobe ( Fig. 2 E), and a rounded apex of the tergal spur ( Fig. 2 C). Mycomya niltava also differs from M. sachak on the basis of the same two characteristics. The base of the wide tergal comb of M. niltava is widely setose ( Fig. 3 A), whereas its medial part is widely bare in M. sachak sp. n. and M. goral sp. n. The hypopygium of M. niltava generally resembles also that of M. sinica Yang & Wu, 1989, but differs from it e.g. in the shape of the tergal lateral appendage and that of the gonostylus. The tergal lateral appendage of M. sinica is broad and apically pointed and its gonostylus has a long membranous lobe.

Etymology. The species name is a noun in apposition and refers to the bird genus Niltava Hodgson in the family of Old World flycatchers, Muscicapidae .

Mycomya sachak sp. n. Figs. 4 A–F

Material studied. Holotype. ♂. NEPAL, Bogara, 1700–2200 m, 83°23'E. 28°35'N. Malaise trap Sept. 20–21. 1971 A. Nakanishi (in KUC). Same data as holotype, but 19–20. IX. 1971, 2 ♂♂ (in KUC); same data, but 21–22. IX. 1971, 1 ♂ (in KUC); same data but not by Malaise trap, 21.IX. 1971, 3 ♂♂, 9 ♀♀ (in KUC).

Description. ♂. Head. Palp, other mouthparts and face yellow, posterior parts of head brown. Antenna brownish, scape, pedicel and base of 1st flagellomere yellow. Length of 1st flagellomere about 3.5x, 2nd flagellomere 2x, its width. Thorax. Pronotum yellow, with 3–4 long setae. Scutum yellowish with 3 brown, fused, longitudinal stripes. Anepisternum yellowish to brownish. Preepisternum yellowish, ventral half brownish. Scutellum brownish, with 4 long setae. Laterotergite brownish. Mediotergite brownish, bare. Wing. Length 3.3–3.7 mm. Wing hyaline. Sc ending in R1 near middle of small cell, Sc1 missing. Apical part of Sc bearing 7–13 small setae. Small cell 1– 2 x as long as wide. Cu fork distal to M fork. M ratios: 0.94–1.17, 1.26.–1.63. Cu ratios: 0.84– 0.85, 1.37–1.45. Small setae: M petiole: 0; M1: 0; M2: 0; Cu petiole: 0; Cu1: 0; Cu2: 0. Halter pale yellowish, apex brownish. Legs. Coxae and femora yellow, tibiae and tarsi brownish to brown. Coxa 2 without spur. Leg ratios: bt1:t1 = 0.71–0.77, bt2:t2 = 0.64–0.66, bt3:t3 = 0.57–0.59. Abdomen. Brownish. Hypopygium. Figs. 4 A–F: yellowish. Tergal part with 1 wide outer comb of spines and 2 very narrow inner combs, both with about 2 spines.

Only lateral parts of base of outer comb with some setae, median part widely bare. Dark inner spur of tergal part apically slightly rounded ( Fig. 4 C). Tergal lateral appendage setose, relatively short, slender, less than 4x as long as wide, only slightly curved, with some flattened, curved apical-subapical setae ( Fig. 4 D). Gonostylus straight, more than 4x as long as wide, with 2 teeth and large membranous lateral lobe slightly shorter than main branch of gonostylus ( Fig. 4 E). Sternal submedian filament very long, curved, extending far beyond apex of aedeagus. Aedeagus with some subapical curved setae on inner side ( Fig. 4 F). Female. Wing length 3.7–4.0 mm. Thorax as in ♂. Abdomen brownish, sternites 1–5 yellowish. Leg ratios: bt1:t1 = 0.77–0.78, bt2:t2 = 0.66–0.70, bt3:t3 = 0.61–0.62. Terminalia yellow.

Discussion. Mycomya sachak resembles M. goral sp. n., but differs from it in its slightly larger size and in the absence of the isolated strong flattened seta in the middle of the inner margin of the tergal lateral appendage. The lateral appendage has several apical and subapical flattened curved setae close to each other ( Fig. 4 D). The tergal lateral appendage of M. sachak is wider than M. goral ( Fig. 2 D), and its subapical setae are shorter than the basal setae, whereas there is no clear difference in their respective lengths in M. goral .

Etymology. The species name is a noun in apposition and refers to the name of the snow leopard in the Tibetan Bhotia language.

Figures

FIGURE 2. Mycomya goral sp. n., male hypopygium. A – B tergal and sternal views, C apex of tergal spur, lateral view, twice enlarged, D tergal lateral appendage, twice enlarged, E gonostylus, twice enlarged, F apex of aedeagus, sternal view, twice enlarged. Scale 0.2 mm.

FIGURE 3. Mycomya niltava sp. n., male hypopygium. A – C tergal, sternal and lateral views, D apex of tergal spur, lateral view, twice enlarged, E tergal lateral appendage, twice enlarged, F gonostylus, twice enlarged, G tergite 8. Scale 0.2 mm.

FIGURE 4. Mycomya sachak sp. n., male hypopygium. A – B tergal and lateral views, C apex of tergal spur, lateral view, twice enlarged, D tergal lateral appendage, twice enlarged, E gonostylus, twice enlarged, F apex of aedeagus, lateral view, twice enlarged. Scale 0.2 mm.