Morellia (M.) quadriremis van Emden, 1965

Pamplona, Denise, Nihei, Silvio S., Couri, Márcia S. & Pont, Adrian C., 2016, Taxonomy of Morellia Robineau-Desvoidy (Diptera: Muscidae): revision of the subgenera Morellia s. str. and Parapyrellia Townsend, Zootaxa 4163 (1), pp. 1-110 : 77-78

34. Morellia (M.) quadriremis van Emden, 1965

( Figs 135–138 )

Morellia quadriremis van Emden, 1965: 105 . Type locality: SRI LANKA, Trincomalee Hot Wells. Holotype Ƌ, BMNH (seen).

Redescription. Male. Colour. Black, with grey pruinosity. Frontal vitta black; face, parafacial and gena black with silver pruinosity; antenna black with grey pruinosity; palpus brown; mesonotum metallic black with a silver-dusted central vitta along presutural acrostichals, pleura grey pruinose; legs brown; haltere yellow; calypters whitish, upper calypter on outer half white and opaque; wing without dark markings; abdomen with a shifting tessellated pattern and with silver pruinosity.

Body length: 6.0 mm; wing length: 5.0 mm.

Head. Eyes bare but with the usual minute and scattered hairs, separated at level of anterior ocellus by 0.10– 0.11 of head width; frons divergent towards lunule; ocellar triangle with 3 pairs of setae; about 13 pairs of weak frontal setae; postpedicel 1.7–1.9 times the length of pedicel; pedicel with 2 setae; arista plumose, with 9–11 cilia, bare apicad, and with some secondary plumes as far as level of 2nd to 4th cilia; vibrissa shorter than arista; 1 strong supravibrissal seta, half as long as vibrissa, and few setulae that extend to level of middle of postpedicel; 5– 7 subvibrissal setae followed by 6–11 genal setae.

Thorax- Acrostichals 0+2, the first pair weaker; dorsocentrals 0+2; intra-alars 1+1; presutural 1; prealar 1, shorter than half the length of first supra-alar; supra-alars 2; postpronotals 3; notopleuron with 2 long, subequal setae and covered with ground-setulae; postalars 2; intrapostalar 1; scutellum with 1 basal, 2 discal and 1 apical pairs of setae, and latero-ventrally with many setae, the basalmost ones downcurved; postalar wall bare; suprasquamal ridge with short pile; proepisternal depression bare; proepimeron with about 4 long setae and many setulae; prosternum broad and bare; anepisternum with a posterior row of about 7 setae, with 4–5 weak setae upcurved on upper-anterior corner, and covered with long setulae; katepisternals 1+2; anepimeron extensively setulose; katepimeron with 2 long setulae; anatergite bare; katatergite bare. Legs: fore femur with a posterodorsal row of setae which are shorter basad, a posteroventral row of setae on basal two-thirds, a ventral row of setae, 4 spines at apical third, and a ventral excavation at apex; fore tibia with an anterodorsal row of short and strong setae, many ventral setulae on apical half, 1 dorsal and 1 anteroventral preapical setae, and 1 ventral apical seta; fore tarsus with 2 long setae on basal segment; mid femur with a tubercle at apex of anterior to posterodorsal surfaces bearing about 8 short spines, 7 ventral setae on basal half, and an anterodorsal row of setae on basal half; mid tibia with an anterodorsal comb-like ctenidium of very short spines on basal half, 4 short and strong posterior setae on basal two-thirds, 2 posteroventral setae on middle third and 1 strong seta on apical third, 1 anterodorsal and 1 dorsal preapical setae, and 1 anteroventral, 1 posteroventral and 1 ventral apical setae; hind femur with an anteroventral row of setae, strongest on apical third, an anterodorsal row of setae, 3 dorsal setae at base, a ventral to posteroventral row of setulae on basal half, shorter but stronger in the middle, 2 posterodorsal preapical setae, a group of posterior setulae at base and few posteroventral setulae on apical fourth; hind tibia with 2 short and subequal anteroventral setae on middle third and other longer setae on apical third, an anterodorsal row of short setae on basal two-thirds of which one at middle third is longer, a dorsal row of short but strong setulae, a long and strong posterodorsal seta on apical third, 1 dorsal and 1 anterodorsal preapical setae, 1 anteroventral and 1 posteroventral apical setae, and a posterior brush-like ctenidium; hind tarsus with a golden brush ventrally. Wing with a non-uniform covering of microtrichia; dm-cu almost straight; stem-vein bare on dorsal surface, and ventrally with about 4 short setulae before humeral cross-vein and bare after humeral cross-vein; R2+3 bare on dorsal and ventral surfaces; R4+5 setulose from base to midway to r-m on dorsal surface.

Abdomen. With a lateral tuft of setulae on syntergite 1+2; tergites 3–5 with a few setae laterally; tergite 5 with apical setae; sternite 1 setulose; posterior margin of sternite 5 concave and with setae, basal half bare ( Fig. 135 ).

Terminalia. Cercal plate ventrally with 3 spines; surstylus with about 4 setulae basally and 2 setae apically ( Figs 136–137 ); aedeagal apodeme narrowed apically; basiphallus long; aedeagus bare; gonopod with one short seta and 4 long and strong setae on basal half, and one short seta on apical half; paramere with many setae apically ( Fig. 138 ).

Female. Unknown.

Material examined. SRI LANKA: Trincomalee Hot Wells , 7.ix.1890, J.W. Yerbury, holotype Ƌ ( BMNH) . INDIA: Bihar, 20.x.1913, 1 Ƌ paratype ( BMNH) .

Distribution. ORIENTAL: India (several states), Sri Lanka.

Figures

FIGURES 131 – 143. Males of Morellia (M.) spp. 131 – 134. Morellia (M.) pseudonigrisquama Shinonaga & Tumrasvin. 131. Sternite 5. 132. Cercal plate. 133. Left surstylus. 134. Phallic complex, lateral view. 135 – 138. Morellia (M.) quadriremis van Emden. 135. Sternite 5. 136. Cercal plate. 137. Left surstylus. 138. Phallic complex, lateral view. 139 – 143. Morellia (M.) setosa Zielke. 139. Sternite 5. 140. Cercal plate. 141. Cercal plate, ventral view. 142. Left surstylus. 143. Phallic complex, lateral view.