Heterometrus spinifer (Ehrenberg, 1828), 1906

Prendini, Lorenzo & Loria, Stephanie F., 2020, Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005, Bulletin of the American Museum of Natural History 2020 (442), pp. 1-480 : 290

Heterometrus spinifer (Ehrenberg, 1828)

Figures 10 , 23C, D , 37C, D , 50E–H , 67E , 68E ,

Figures

FIGURE 10. Phylogeny of Asian forest scorpions (Scorpionidae: Heterometrinae Simon, 1879, stat. nov.), obtained by simultaneous phylogenetic analysis of 186 morphological characters and 4188 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci (Loria and Prendini, in press). Maximum likelihood tree with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles parallel derivations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (appendix 1).

FIGURE 23. Heterometrus Ehrenberg, 1828, carapace, dorsal aspect. A, B. Heterometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev., A. ♂ (AMNH), Thale Ban, Thailand, B. ♀ (AMNH), Khao Pu-Khao Ya, Thailand. C, D. Heterometrus spinifer (Ehrenberg, 1828), C. ♂ (AMNH), D. ♀ (AMNH), Cameron Highlands, Malaysia. Scale bars = 2 mm.

FIGURE 37. Heterometrus Ehrenberg, 1828, sternum and pectines, ventral aspect. A, B. Heterometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev., A. ♂ (AMNH), Thale Ban, Thailand, B. ♀ (AMNH), Khao Pu- Khao Ya National Park, Thailand. C, D. Heterometrus spinifer (Ehrenberg, 1828), C. ♂, D. ♀ (AMNH), Cameron Highlands, Malaysia. Scale bars = 2 mm.

FIGURE 50. Heterometrus Ehrenberg, 1828, legs I–IV, tibiae, basitarsi and telotarsi, ventral aspect. A–D. Heterometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev., ♂ (AMNH), Thale Ban, Thailand. E–H. Heterometrus spinifer (Ehrenberg, 1828), ♂ (AMNH), Cameron Highlands, Malaysia. Scale bars = 2 mm.

FIGURE 67. A, B. Gigantometrus Couzijn, 1978, stat. nov., and C–E. Heterometrus Ehrenberg, 1828, metasoma and telson, dorsal aspect. A. Gigantometrus swammerdami (Simon, 1872), comb. nov., lectotype ♂ (Palamnaeus swammerdami flavimanus Pocock, 1900, syn. nov.) (BMNH 1899.11.2.292-3). B. Gigantometrus titanicus (Couzijn, 1981), comb. nov. et stat. rev., ♂ (AMNH), Polonnaruwa, Sri Lanka. C. Heterometrus glaucus (Thorell, 1876), comb. nov. et stat. rev., ♀ (AMNH), Nias Island, Indonesia. D. Heterometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev., ♂ (AMNH), Thale Ban, Thailand. E. Heterometrus spinifer (Ehrenberg, 1828), ♂ (AMNH), Cameron Highlands, Malaysia. Scale bars = 5 mm.

FIGURE 68. A, B. Gigantometrus Couzijn, 1978, stat. nov., and C–E. Heterometrus Ehrenberg, 1828, metasoma and telson, lateral aspect. A. Gigantometrus swammerdami (Simon, 1872), comb. nov., lectotype ♂ (Palamnaeus swammerdami flavimanus Pocock, 1900, syn. nov.) (BMNH 1899.11.2.292-3). B. Gigantometrus titanicus (Couzijn, 1981), comb. nov. et stat. rev., ♂ (AMNH), Polonnaruwa, Sri Lanka. C. Heterometrus glaucus (Thorell, 1876), comb. nov. et stat. rev., ♀ (AMNH), Nias Island, Indonesia. D. Heterometrus laevigatus (Thorell, 1876), comb. nov. et stat. rev., ♂ (AMNH), Thale Ban, Thailand. E. Heterometrus spinifer (Ehrenberg, 1828), ♂ (AMNH), Cameron Highlands, Malaysia. Scale bars = 5 mm.