Pterostichus pinguedineus (Eschscholtz, 1823)

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 548-549

Pterostichus pinguedineus (Eschscholtz, 1823)

Poecilus pinguedineus Eschscholtz, 1823: 106. Type locality: "Unalaschka [Aleutian Islands, Alaska]" (original citation). Syntype(s) probably in ZILR. Note. This species was also made available the same year, under the name Poecilus pinguedinis , by an illustration in Fischer von Waldheim (1823: plate 19, figure 7).

Feronia frigida Dejean, 1828: 334. Type locality: "Kamschatka [Russia]" (original citation). Lectotype (♀), designated by Ball (1966a: 70), in MHNP. Synonymy established by Ball (1966a: 70).

Argutor mandibularis Kirby, 1837: 31. Type locality: "Lat. 54° [= along North Saskatchewan River]" (original citation). Lectotype [as type] (♂), designated by Ball (1966a: 70), in BMNH. Synonymy established by Ball (1966a: 70).

Feronia subsinuosa Chaudoir, 1868b: 339. Type locality: "Kadjak [Island, Alaska]" (original citation). Holotype [by monotypy] (♀) in MHNP. Synonymy established by Ball (1966a: 70).

Feronia diplogma Chaudoir, 1868b: 340. Type locality: «côte nord-ouest de l’Amérique du Nord" (original citation). Holotype [by monotypy] (♀) in MHNP. Synonymy established by Ball (1966a: 70).

Feronia stuxbergi Mäklin, 1878: 17. Type locality: "Krestowskoj [Taimyr Autonomous Okrug, Siberia, Russia]" (lectotype label). Lectotype (♂), designated by Ball (1966a: 70), in ZMH. Synonymy established by Ball (1966a: 70). Etymology. The specific name honors the Swedish explorer and naturalist Anton Julius Stuxberg [1849-1902] who took part to the Arctic zoological expeditions with Adolf Erika Nordenskiöld in 1875-1876 and 1878-1879. Stuxberg worked at the geological museum in Göteborg until his death.

Feronia despecta J.R. Sahlberg, 1885a: 14. Type locality: "Pitlekaj [Chukchi Peninsula, Russia]" (lectotype label). Lectotype (♀), designated by Ball (1966a: 70), in NRSS. Synonymy established by Ball (1966a: 70).

Feronia sulcipennis J.R. Sahlberg, 1885a: 15. Type locality: "Pitlekaj [Chukchi Peninsula, Russia]" (original citation). Holotype [by monotypy] (♀) in NRSS. Synonymy established by Ball (1966a: 71).

Feronia laeviuscula J.R. Sahlberg, 1885a: 16 [secondary homonym of Pterostichus longicollis var. laeviusculus Letzner, 1852]. Type locality: "Nunamo, S[ain]t Lawrence Bay [Chukchi Peninsula, Siberia, Russia]" (original citation). Holotype [by monotypy] (♀) in NRSS. Synonymy established by Ball (1966a: 71).

Feronia splendida J.R. Sahlberg, 1885b: 49. Type locality: Port Clarence, Alaska (inferred from title of the paper). Holotype [by monotypy] (♀) in NRSS. Synonymy established by Ball (1966a: 71).

Cryobius subnitidulus Poppius, 1906b: 62. Type locality: "Insel Kadjak [Alaska]" (original citation). Syntype(s) [2 originally cited] in ZMH (Poppius 1906b: 64). Synonymy established, under the name Pterostichus subsinuosus (Chaudoir), by Poppius (1908a: 2).

Cryobius holmbergi Poppius, 1906b: 70. Type locality: "Insel Kadjak [Alaska]" (original citation). Holotype [by monotypy] (♀) in ZMH. Synonymy established by Ball (1966a: 71).

Cryobius incognitus Poppius, 1906b: 91. Type locality: "Amer[ica] ross[ica]" (original citation). Holotype [by monotypy] (♀) in ZILR. Synonymy established by Ball (1966a: 71).

Cryobius stuxbergi var. fortestriatus Poppius, 1906b: 103. Type locality: "Briochowskiostroff, Fl[umen] Jenissej, arktisch Sibirien [Russia]" (original citation). Holotype [by monotypy] (♀) in ZMH. Synonymy established by Ball (1966a: 71).

Cryobius stuxbergi repandus Poppius, 1906b: 104. Type locality: "Lena infer[ior] [Yakutia, Siberia, Russia]" (lectotype label). Lectotype (♀), designated by Ball (1966a: 71), in ZMH. Synonymy established by Ball (1966a: 71).

Cryobius alaskensis Poppius, 1906b: 116. Type locality: "Insel Kadjak, Alaska" (original citation). Syntype(s) [4 originally cited] in ZILR. Synonymy established by Ball (1966a: 71).

Pterostichus montanellus Poppius, 1907: 21. Type locality: "White Mountains [New Hampshire]" (original citation). Two syntypes [2 originally cited] in ZMHB (Lindroth 1966: 518) and ZMH (Silfverberg 1987: 20). Synonymy established by Ball (1966a: 71).

Pterostichus beringensis Poppius, 1908b: 4. Type locality: "St. Lawrence Bai [Chukchi Peninsula, Russia]" (original citation). Holotype [by monotypy] location unknown. Synonymy established by Bousquet (2003d: 492).

Cryobius beringi Casey, 1918: 374. Type locality: "S[ain]t Paul Island, Alaska" (original citation). Lectotype [as holotype] (♀), designated by Ball (1966a: 71), in USNM [# 47078]. Synonymy established by Ball (1966a: 71).

Cryobius washingtoni Casey, 1920: 190. Type locality: "White M[oun]t[ain]s, New Hampshire" (original citation). Lectotype (♀), designated by Ball (1966a: 71), in USNM [# 47075]. Synonymy established by Ball (1966a: 71).

Pterostichus anadyricus Csiki, 1930: 651. Replacement name for Pterostichus laeviusculus (Sahlberg, 1885).

Distribution.

This Holarctic species ranges from the Yenisei River in Siberia eastwards to the Mackenzie River delta in western Northwest Territories; isolated on the top of some mountains in New England and on the Shickshock Mountains of the Gaspé Peninsula, Quebec [see Ball 1963: Fig. 5]. The record from Wyoming (Lavigne 1977: 47) is in error. Fossil remnants of this species, dated between about 12,000 and 18,100 years B.P., have been unearthed in central and southeastern Iowa (Schwert 1992: 77; Baker et al. 1986: 96) and northeastern Pennsylvania (Barnosky et al. 1988: 178); others, older than 33,000 years B.P., have been found in southwestern Ontario (Warner et al. 1988: 37).

Records.

CAN: NT, QC, YT USA: AK, ME, NH, VT - Holarctic

Note.

Platysma poppiusianum Jacobson, 1907 (a replacement name for Pterostichus insulicola Poppius, 1906) was listed as a junior synonym of this species by Ball (1966a: 71), as a junior synonym of Pterostichus subgibbus (Motschulsky, 1860) by Kryzhanovskij et al. (1975: 137), and treated as a valid species by Eremin (1990) and Kryzhanovskijet al. (1995: 102).