Salcedia schoutedeni Alluaud, 1930

Balkenohl, Michael, 2020, A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini), ZooKeys 901, pp. 1-81 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.901.39432
persistent identifier	https://treatment.plazi.org/id/5DEE114B-7FB4-5465-93D6-7C204D8440B6
treatment provided by	ZooKeys by Pensoft
scientific name	Salcedia schoutedeni Alluaud, 1930
status

Treatment

Salcedia schoutedeni Alluaud, 1930 Figs 14 View Figures 14–19 , 32 View Figures 26–35 , 48 View Figures 44–51 , 58 View Figures 58–63 , 73 View Figures 68–76 , 84 View Figure 84

Salcedia schoutedeni Alluaud, 1930: 21; Alluaud 1932: 2; Csiki 1933: 641; Alluaud 1935: 19; Burgeon 1935: 159; Andrewes 1936: 64.

Salcedia schoutedeni schoutedeni Alluaud, 1930: Reichardt 1975: 103; Dostal 1993: 121; Lorenz 2005: 155.

Type material.

Holotype: ♀, with labels and data: white, printed and handwritten “MUSÉE DU CONGO Kwamouth 9-VII-1921 Dr H.Schouteden" / white, printed black framed "R. DÉT. 1783 c" red, black double framed “HOLOTYPE” / white, black framed, handwritten and printed " Salcedia Schoutedeni Type Alluaud, det. 1930" (MRACT).

Remark.

In the holotype both of the front legs are broken and glued.

Additional material.

1 ♂, 1 ♀, 22 specs., Léopoldville I-1947 Dr E. Dartevelle COLL. MUS. CONGO (MRACT, CBB); 1 ♀, same data (MNHN); 1 spec., Léopoldville -1930 Eg. Devroye MUSÉE DU CONGO / R. DÉT. 2019 b / Salcedia schoutedeni All. Alluaud. det. 1931 (MRACT); 1 spec., Léopoldville -1930 E. Devroey MUSÉE DU CONGO (MRACT); 1 spec., same data but -1931 (MRACT).

Diagnosis.

A large sized species, with sub-elongate outline of the elytra with maximum width at middle and the pronotum with a rudiment of the outer lateral carina. The pseudohumerus is rectangular with a distinctly laterally projecting tooth. The antennomeres are elongate. Distinguished most clearly from the similar species S. nigeriensis by the pronotum with the lateral margin converging from middle to the anterior angles, the pseudohumerus of the elytron with a distinct tooth, and the female coxostylus with six large setae and one SSO. In addition, S. schoutedeni has a distinct keel on the frons of the head.

Redescription.

Measurements in Table 1 View Table 1 .

Colour and surface: Anthracite grey, shiny; top of carinae on head, pronotum and elytra as well as margins of pronotum and elytra opaque, covered with pale grey pili; legs piceous, mandibles, antennae and palpi piceous.

Head: Less than two-thirds of width of pronotum. Outline semi-circular. Clypeus wide, convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by distinct notches, with slightly raised pentagon-shaped field at middle, separated from frons by deep transverse furrow. Frons with two raised paramedian carinae, joining anteriorly into a V-like keel, with two small glossy teeth bilaterally anterior to central keel, with two indistinct short parallel carinae paralaterally at base; frons separated from supraorbital plates by deep broad furrows; with conspicuously deep and broad pit at front-eye level; supra-antennal and supraorbital plates margined, margin of supraorbital plate distinctly raised, carina-like, supra-antennal plates vaulted. Base with narrow emargination at middle, sharply rectangular laterally (angle around 109°). Eyes large, convex, genae slightly convex, both of them clearly visible from above, with nearly circular shape in lateral view. Antenna with segments five to ten elongate (L/W 1.25), densely pubescent, segments two to four scarcely pubescent, scapus with irregular reticulation. Labrum nearly straight anteriorly. Mandible moderately short, wide, slightly arcuate at apex. Mentum small, with slight tooth at middle, epilobes wide, projecting and angled anteriorly, margined anteriorly, surface with small pits anteriorly, with isodiametric reticulation basally.

Pronotum (Fig. 32 View Figures 26–35 ): Outline rectangular, transverse, two-fifth wider than long. Lateral margin convex, distinctly converging anteriorly, maximum width behind middle. Lateral margin crenulated, with 12-13 small tubercles, tubercles larger basally, with two notches at posterior angles. Base straight laterally, with declining flat keel at middle pointing posteriorly. Disc flattened in lateral view, with two raised paramedian carinae parallel to median line and diverging posteriorly, with median line small, long, and ending in pits anteriorly and posteriorly, with two additional shorter carinae bilaterally, the paralateral one joining with the paramedian carinae at base and forming tooth-like tubercle pointing posteriorly, without anterior extension, without inner lateral carina, with very small outer lateral carinae. All carinae sub-crenulate. Lateral margin broadly wing-like bent up, with six large and deep transverse pits, the basal and/or frontal ones partly separated into two smaller pits. Space between carinae and pits smooth.

Elytron (Fig. 48 View Figures 44–51 ): Flattened in anterior half (lateral view), convex in frontal view. Sub-elongate, margin straight in anterior half up to pseudohumerus, moderately convex to apex, maximum width at middle, sub-undulate. Pseudohumerus rectangular, tooth distinctly projecting laterally. Apex rounded, with small but acute tooth at suture. Disc with interneur six sub-crenulate, interneur three slightly carinate in basal third, interneur two running up to apex as slightly convex line, conspicuously raised; interneur four running in parallel to interneur six, almost reaching base, shortened at apex. Interneur five and six with two rows of serial pits partly merging transversally.

Hind wings: Fully developed.

Lower surface: Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with one row of slightly transverse pits, lateral margin of elytron smooth. Metepisternum elongate, with broad longitudinal groove. Metasternum, abdominal sternites one and five with numerous irregularly situated larger and smaller pits, sternite two to three smooth, with band of small punctures at middle. Last three abdominal sternites laterally with isodiametric reticulation, sternite one with longitudinal reticulation, two with transverse reticulation. Sternum four to six slightly sulcate. Sternum six at apical margin with hollowed out transverse impression at apex.

Legs: Profemora with surface indistinctly reticulated. Protibia with short, robust, moderately curved terminal spine, laterally with laterally with two larger and two smaller teeth, dorsally and ventrally with two carinae. First tarsomere distinctly elongated, as long as tarsomeres two to five together, front and intermediate legs with tarsomeres two to four broadened and somewhat flattened.

External sexual dimorphism: Sternum six with the hollowed out transverse impression at the apical margin is obtusely longitudinally sulcate in females.

Male genitalia (Fig. 58 View Figures 58–63 ): Median lobe stout, in dorsal view distinctly and regularly arcuate, straight to apex, in lateral view straight, apparently no fine pili visible, apical part bent ventrally apex dorsally, in cross section oval at apex. Oroficium small. Endophallus with group of longer microtrichia near oroficium, with additional longer group of brush-like microtrichia basally. Dorsal paramere relatively short, bisinuate, with short apophyses; ventral one shaped like a convex spatula, with fine seta at apex; both parameres slightly distorted.

Female genitalia (Fig. 73 View Figures 68–76 ): Coxostylus regularly broadened to base, distinctly curved, acute at apex, with indistinct carina dorsally in apical third, at end of basal third with one strong and five slender nematiform setae laterally, SSO with one fine pilus.

Variation: The tubercles at the lateral margin of the pronotum are more or less strongly developed. Number and shape of the pits on the ventral surface varies to a small degree.

Distribution

(Fig. 84 View Figure 84 ). Known from several localities in the west of the Democratic Republic of the Congo along the lower course of the river Congo. The locality Kwamouth refers to the short estuary of the river Kasai into the River Congo. The location “Zambèze, Nova Chupanga près Chemba" quoted in Alluaud (1935) as new record for S. schoutedeni refers to Salcedia africana .