Stolonis catenarius Will

Will, Kipling W., 2005, The Neotropical genera Oxycrepis Reiche and Stolonis Motschulsky: a taxonomic review, key to the described species and description of new Stolonis species from Ecuador (Coleoptera: Carabidae: Loxandrini), Zootaxa 1049, pp. 1-17 : 15-16

Stolonis catenarius Will , new species

( Figs. 2 E, 3Q–T)

Type Material. HOLOTYPE: ɗ ( EMEC), genitalia dissected. Labeled: “ ECUADOR, Napo Prov., Yasuni Scientific Station, 22:IV: 1998, 210m, Col.K.Will, Headlamp"/"U.C. Berkeley EMEC 1003678"/[red label] " Holotype, Stolonis catenarius, K.Will ". ALLOTYPE: Ψ ( EMEC), Labeled same as holotype /"U.C. Berkeley EMEC 1003678"/[red label] "Allotype, Stolonis catenarius, K.Will ".

PARATYPES: Ecuador: Orellana Prov.: Yasuni Scientific Station, 00°40'36"S 76°24'02: 1ɗ ( QCAZ), " 210m, Col.K.Will, Headlamp" EMEC 1003674; 2 m EMEC 1003679 ( EMEC), EMEC 1003681 ( USNM); 1Ψ, ( QZAC), 21:iv:1998, EMEC 1003673; 1Ψ, ( EMEC), 25:iv: 1998, 210m, Col.K.Will, heading in palm swamp, EMEC 1003675; 1Ψ( EMEC); 25:iv: 1998, 210m, Col.K.Will, headlamp, EMEC 1003676.

Etymology. Specific epithet is based on the Latin word catenarius , a, um, "chains" and refers to the series of closely set punctures of the elytral striae.

Diagnosis. Differs from other Stolonis species by the combination of broad pronotum ( Fig. 2 E) with broadly expanded lateral margins, immaculate elytra and evenly and densely punctate elytral striae.

Description. Length 6.1mm (6.1–6.0mm). Dorsal surface black, vaguely paler at apex, shiny, lightly iridescent; antennomeres 1–3 brunneous or palpi paler; anternomeres 4–6 and 11 black, 7–10 white; mouth parts brunneous and legs flavotestaceous with coxae and trochanters concolorous with ventral surface or slightly more infuscated.

Form of head slightly broader than average build; eyes only slightly prominent; not or very slightly constricted behind eyes; frontal impressions well marked but not clearly delimited medially, shallow, nearly punctiform, short, length much less than half distance from base of clypeus to anterior supraorbital setae; frons between eyes shiny, scarcely iridescent near base, microlines not evident.

Pronotum ( Fig. 2 E); clearly broader than long; narrowly constricted at base; anterior submarginal sulcus deep and complete; basal impressions short, 1/3 length of pronotum or less, very shallow apically and basally, apically divergent when evident; lateral margins broadly rounded and widely explanate, constricted just apicad hind setae; not evident basad hind setae; without angular process at the hind setae; shallowly punctate over base and in basal impressions. Elytra, elongate oval, length 3.8mm, overall width 2.3mm; slightly convex, rounded and slightly sinuate near apex; humeri sloped and rounded; striae with moderately deep small, dense, punctures, somewhat less evident apically; intervals, broad, slightly convex, only slightly crenulate. Legs, slender; meso­ and metatarsi with prominent external sulcus; fifth tarsomeres ventrally with two pair of very small setae or glabrous. Ventral surface shiny, clearly iridescent; mesosternum with 6–10 deep, broad punctures; metasternum laterally with 4–8 broad, more or less deep punctures; metepisternum with medial sulcus deeply impressed, slightly punctate, slightly longer than wide (l/w= 1.2).

Base of abdominal sternum II with row of dense, moderately deeply impressed punctures; base of sterna IV–VII with very narrow, dull band of coarse irregular microsculpture, restricted to lateral region in some specimens; sterna IV–VI with one pair paramedial setae; in male one pair paramedial setae on VII, female with two pairs. Aedeagus ( Figs. 3 Q–T) with median lobe bluntly rounded, slightly asymmetrical tip, ostium dorsal and large; ventral surface well sclerotized; in repose endophallus with clearly defined scale field left of midline from near apex to almost 1/2 length of median lobe, best visible in ventral view. Seven other spines arranged as in figure 3R. Female reproductive tract with large right dorsolateral bursal pouch, spermatheca broadly connected and stout; appended gland duct 3.5× length of spermatheca attached to spermatheca base; laterotergites IX with scattered setae; gonocoxite­1 with 5–6 apical setae; gonocoxite 2 with 2–3 lateral ensiform setae, 2 apical nematiform setae. Pygidial gland form simple, without any additional lobes, chemical compounds produced unknown.

Natural history and collecting information. Night active in very wet areas of primary forest. Taken treading vegetation in muddy areas and near fallen Ficus and palm plants. Collected with S. yasuni and S. spinosus .

Figures

FIGURE 2. Pronota of: A— Stolonis tapiai, B— Stolonis scortensis, C— Stolonis spinosus, D— Stolonis yasuni, E— Stolonis catenarius.

FIGURE 3. Aedeagus dorsal, left lateral, right lateral and ventral views of: A – D— Stolonis tapiai, E – H— Stolonis scortensis, I – L— Stolonis spinosus, M – P— Stolonis yasuni, Q – T— Stolonis catenarius.