Anthracus javaensis, Jaeger, 2015

Jaeger, Bernd, 2015, Revision of the maculate species of the Anthracus annamensis group from the East Palaearctic and Oriental Regions. Part 2. A redescription of Anthracus nesophilus (ANDREWES, 1936) and six new species from Nepal, India and SE Asia (Coleoptera, Carabidae, Harpalini, Stenolophina), Linzer biologische Beiträge 47 (2), pp. 1361-1396 : 1373-1374

publication ID	https://doi.org/ 10.5281/zenodo.5193661
DOI	https://doi.org/10.5281/zenodo.6546858
persistent identifier	https://treatment.plazi.org/id/5C580E35-A81D-F11A-FF5E-FEC4FE699D28
treatment provided by	Felipe
scientific name	Anthracus javaensis
status	sp. nov.

Treatment

Anthracus javaensis View in CoL nov.sp.

( Figs 19-20 View Figs 19-20 , 62 View Figs 62-67 , 68 View Figs 68-73 , 80 View Fig 80 )

Type material: Holotype: ♂ ( NHMW) labelled " INDONESIA: W Java / Situ Lembang, ca. 1500m / 15 km N Bandung / lg. Schuh 7.8.1994 " and "HOLOTYPE ♂ / Anthracus / javaensis sp. n. / des. B. Jaeger 2015". [red label]

Etymology: The species name refers to the known distribution of the species.

Description: General appearance as figured ( Fig. 19 View Figs 19-20 ). Body length 4.0 mm; width 1.55 mm.

Shiny, pronotum weakly, elytra moderately iridescent. Head dark brown, with clypeus reddish brown, labrum and mandibles reddish yellow (inner margins and apices of the latter blackish). Pronotum darker than head, dark to blackish brown, with lateral furrows paler in basal half. Elytra dark reddish yellow, with each elytron having a very large almost blackish central macula, expanding laterally to interval 8, and leaving base, apex and first interval reddish yellow. Legs and palpi pale yellowish brown, antennae with first two antennomeres yellowish brown, remaining ones moderately infuscated. Ventral surface mainly brown to reddish brown, with proepisterna markedly darker brown, metepisterna and lateral parts of abdominal sternites moderately darker brown, epipleura reddish brown.

Head ( Figs 19-20 View Figs 19-20 ) including eyes 0.86 times as wide as pronotum, with eyes moderately prominent (head 1.61 times as wide as head between eyes). Labrum with apical margin linear but somewhat sloped down to the right side. Mandibles medium sized, left mandible rather short and somewhat obtuse at apex, but not thickened or truncate. Antennae moderately long, 2.46 times as long as pronotum and 0.86 times as long as elytra. Microsculpture on clypeus and labrum distinct and almost isodiametric, partly weakly transverse, on frons with very lightly impressed (partly obliterated) and on vertex with moderately impressed isodiametric meshes becoming weakly transverse in front of pronotal apical margin.

Pronotum ( Figs 19-20 View Figs 19-20 ) 1.30 times as wide as long, 1.27 times as wide as head, widest in second quarter, lateral seta inserted a little posterior to beginning of second quarter. Apical margin almost rectilinear, anterior angles narrowly rounded at tips, weakly projecting forward. Sides convex in anterior half and rectilinearly narrowed to posterior angles, which are obtuse and moderately rounded. Basal margin very weakly convex medially and moderately oblique to posterior angles,. Lateral furrows moderately wide in anterior half becoming only slightly wider in posterior quarter where they are fused with baso-lateral impressions. Baso-lateral impressions medium sized, distinctly delimited from the convex pronotal disc and the weakly depressed medial part of base, flattened to basal and lateral margins. Anterior transverse impression suggested. Microsculpture on disc with moderately impressed moderately transverse meshes, becoming weakly impressed and strongly transverse at sides, at baso-lateral impressions and lateral furrows with distinctly impressed isodiametric to weakly transverse meshes.

Elytra ( Fig. 19 View Figs 19-20 ) rather long, 1.59 times as long as wide, 2.84 times as long and 1.39 times as wide as pronotum. Sides weakly widened posteriad, widest just posterior to middle. Subapical sinuation weak. Elytral striae distinctly impressed and impunctate, intervals rather flat, becoming weakly narrowed and moderately convex at apex. Microsculpture on scutellum and around basal pore isodiametric, on elytral intervals only with traces of very lightly impressed transverse lines.

Metepisterna long, distinctly narrowed posteriad, at inner margin about 1.6 times longer than wide at basal margin. Prosternum medially with 5 and close to apical margin with a row of 8 medium long setae. Prosternal process with 1 distinct seta.

Protarsomeres 1-4 of males markedly and mesotarsomeres 2-4 moderately dilated. Protarsomere 4 markedly and mesotarsomere 4 moderately bilobed. Pro- and mesotarsomeres 1-4 of males with biseriately arranged adhesive hairs on ventral surface.

Median lobe of aedeagus ( Figs 62 View Figs 62-67 , 68 View Figs 68-73 ) large, with general appearance as figured. Apical lamella rather short and broad, not narrowed (dorsal aspect), the apex abruptly bent upward (lateral aspect). Internal structures composed of 3 larger subapical teeth, and three medium sized teeth in basal half.

Remarks and Comparisons: A. javaensis nov.sp., A. siamensis nov.sp. from Laos and Thailand and A. skalei JAEGER from Nepal, NE India, Bhutan and Bangladesh represent a complex of closely related, allopatrically distributed taxa which are characterized by a similar shape of the median lobe of the aedeagus with apex short in dorsal view and abruptly bent dorsad in lateral view. The species of this complex are partially very similar in general appearance, but can be distinguished by constant different internal structures of the aedeagus. A. javaensis nov.sp. is the largest species of this complex and differs from A. siamensis and A. skalei by its large size, usually also by some body proportions (see HW/PW, EW/PW, EL/PL) and the different internal structures of the aedeagus ( Figs. 62-75 View Figs 62-67 View Figs 68-73 View Figs 74-79 ). From A. nesophilus which occurs also in western Java it can be separated by characters given under this species. From A. sumatraensis it differs by the dark head and pronotum, larger size, some body proportions (HW/PW, EL/EW), and the different external shape and internal structures of the median lobe of the aedeagus ( Figs. 62 View Figs 62-67 , 68 View Figs 68-73 , 50-55 View Figs 50-55 ).