Naddia iacobi, De, 2014

De, G., 2014, The genus Naddia in Borneo (Coleoptera: Staphylinidae: Staphylininae), Linzer biologische Beiträge 46 (2), pp. 1737-1765 : 1743-1744

Naddia iacobi nov.sp.

(Figs 2, 2a, 2al)

Holotype: SARAWAK, 2 nd Division , River Skrang , XI.1976, G. de Rougemont (in CRO)

Additional material: 1ex.: SARAWAK,4 th Division , Gn. Mulu NP, pitfall-trap, fish bait, MD forest, 500-800 m, Naddia drescheri Cam. P.M. Hammond det. 1979 (in NHML) ; 1: BRUNEI, ii-iii.1992, E115 7˚ N4 34˚ Kuala Belalong FSC, Ground Malaise trap, N. Mawdsley coll. BMNH{E} 1991-173, Naddia drescheri Cam. det. R.G. Booth 2009 (in NHML) ; 4: INDONESIA, Borneo, Kalimantan Tengah, Busang / Rekut confl. 0˚03’S 113˚59’E, Flight Intercept FIT 7, Brendell / Mendel , August 2001, ‘ Barito Ulu 2001’, BMNH(E) 2001-191 (in NHML) .

Description: Proportions of holotype: length: 12 mm; length of head: 1.8; total length of head: 2.0; breadth of head: 2.15; length of eye: 0.8; length of temple: 0.7; length of antenna: 2.4; length of pronotum: 2.1; breadth of pronotum: 2.1; length of elytron: 2.5; breadth of elytra: 2.7. Habitus: fig. 2.

Temples dilated posteriorly; eyes very large and prominent, their diameter in dorsal view greater than length of temples. Antennomeres 1-3 elongate; 3-4 globose, 7-10 increasingly transverse but not asymmetrical.

Integument of head, pronotum and abdomen black, that of elytra rufo-testaceous; genital segment brown; palpi reddish, antennae fuscous; legs rufo-testaceous. Head and pronotum with a few scattered pale hairs and some dark setae on the sides; pubescence of scutellum very dense, deep black elytra entirely covered with rather dense, pale brassy pubescence, the sides with a few dark setae; pubescence of abdominal tergites black; tergite III in addition with longer, sparse golden pubescence on entire surface; tergite IV with a narrow lateral fasciae of golden pubescence extending the entire length of tergite; tergite V with only a small patch of such pubescence in antero-lateral corners; tergite VII with a pair of large patches of very bright golden pubescence, the hairs radiating outward from antero-median corner. Legs with pale pubescence and setae.

Sculpture of head very coarse, consisting of large umbilicate punctures near posterior margins, the interstices coalescing to form long parallel longitudinal rugae in front of this, so that individual punctures are no longer visible anteriorly; sculpture of pronotum very coarse, consisting of large close umbilicate punctures on entire surface, leaving only fragments of a mid-longitudinal band appearing as narrow shiny callosities; puncturation of elytra consisting of shallow umbilicate punctures, smaller than on head and pronotum, the interstices as or almost as wide as diameter of punctures and wrinkled, so that the surface is not very shiny; puncturation of abdominal tergites fine and dense near anterior margins, coarser and much sparser posteriorly.

Male: sternite VII modified, the sculpture of depression much finer and denser than that of rest of sternite; in the specimen figured the anterior margin of depression lacks the anterior fringe of long setae (probably detached and lost); sternite VIII with a small, very shallow emargination; aedoeagus: figs 2a, 2al, the median lobe in ventral view very different from that of N. germana , longer and narrower; basal sclerite narrowed to an obtuse point at apex; paramere very short, with an apico-median notch.

The specimens from Sarawak and Brunei in the NHML were determined as Naddia drescheri CAMERON, 1937 , a species described on the basis of an unspecified number of females from Java. I compared the Bornean material with the paratype of N. drescheri in the NHML and found them to belong to a different species ( N. iacobi nov.sp.). The two species are very similar, including the size of the eyes (fractionally smaller in N. drescheri , about as long as temples) and sculpture of the fore-body, but in N. drescheri the sculpture and coloured pubescence of the abdominal tergites are different: in N. drescheri the puncturation of all tergites is finer and sparser, the surface therefore more shiny; tergite VII has very strong and evident transverse micro-striation on its entire surface (micro-sculpture reduced to a few scattered micro-punctures in N. iacobi ); the coloured pubescence of tergites is disposed in the same pattern, but is silvery in N. drescheri instead of golden-brassy. These differences may appear to be slight and inconclusive, but I am confident of the validity of the new species, and am convinced that N. drescheri does not occur in Borneo.

The female specimens from Kalimantan here attributed to N. iacobi are a little larger and more finely punctured than the type and the exx. from Sarawak and Brunei; in this respect they are somewhat intermediate between N. iacobi and N. germana nov.sp., but much closer to the former, including in the ratio of length of eyes/temples.

For comparison with N. germana nov.sp. and comment on the distribution of the two species see below, description of N. germana nov.sp.

Derivation of specific epithet: named in honour of James Brooke, first Rajah of Sarawak, who was A.R. Wallace’s host, not only in his country but in his houses, during the naturalist’s first scientific expedition to Borneo in 1854-56.