Naddia, FAUVEL

De, G., 2014, The genus Naddia in Borneo (Coleoptera: Staphylinidae: Staphylininae), Linzer biologische Beiträge 46 (2), pp. 1737-1765 : 1740-1742

Key to the Bornean species of Naddia

(To fully appreciate the patterns of pubescence it is sometimes necessary to shift the angle of illumination of specimens under the microscope to different positions.)

1 Temples convergent towards base; sides of head, elytra and pleurites of the genital segment devoid of large setae in addition to the fine pubescence; penultimate antennomeres as long as wide, not evidently asymmetrical; puncturation finer; posterior 2/3 rd of elytra with a strong brassy reflex, the entire disc bearing long coppery pubescence; length 12-13 mm. Habitus: fig ................................... asetosa nov.sp.

- Temples parallel-sided or dilated towards base: sides of head, elytra and pleurites of the genital segment with several large setae in addition to pubescence; penultimate antennomeres distinctly transverse and usually more or less asymmetrical; puncturation coarse.............................................................................................................3

3 Elytra red or rufo-testaceous...............................................................................................4

- Elytra black, fuscous, or black with a (sometimes faint) metallic tinge..............................6

4 Pronotum and abdominal tergite III black; temples sub-parallel; elytra red.......................5

- Pronotum reddish brown; abdominal tergite III entirely rufo-testaceous; temples dilated towards base; elytra rufo-testaceous; male unknown .......................... germana ab.?

5 Eyes longer than temples seen in dorsal view; puncturation of head and pronotum coarser and sparser, the surface therefore more shiny; temples and antero-lateral areas of pronotum with only a few scattered pale brassy setae. Smaller insect (ca. 12 mm) Habitus: fig. 2; aedoeagus: figs 2a, 2al .......................................................... iacobi nov.sp.

- Eyes shorter than temples seen in dorsal view; puncturation of head and pronotum finer and denser, the surface therefore duller; temples and antero-lateral areas of pronotum with a conspicuous patch of pale brassy setae. Larger insect (ca. 15 mm).Habitus: fig. 3; aedoeagus: fig. 3a, 3al............................................... germana nov.sp.

6 Elytra with a distinct greenish brassy reflex; small species (10-12mm); diameter of eyes in dorsal view greater than length of temples; pronotum with an entire shiny impunctate mid-longitudinal axis. Habitus: fig 4; male tergite VIII (figs 4b, 4bl) with a characteristic modification; aedoeagus: ig. 4a, 4al, the paramere elongate, freely articulated.......................................................................................... aeneipennis CAMERON

- Elytra black or fuscous, sometimes slightly rufescent behind humeral angles, or with a faint bluish tinge; paramere short, adpressed....................................................................11

7 Posterior margins of elytra with a conspicuous dense fringe of pale pubescence; pronotum without trace of an impunctate mid-longitudinal line; tergite VIII entirely reddish-brown; large species (19-22 mm); male sternite VII unmodified. Habitus: fig. 5; aedoeagus: figs 5a, 5al ............................................................................. limbifer nov.sp.

- Posterior margins of elytra with sparse pale and/or dark pubescence, but not forming a conspicuous entire fringe..................................................................................................13

8 Head and pronotum shiny; frons broadly impunctate, very shiny with only scattered smaller punctures, the vertex with an impunctate mid-longitudinal band stretching from impunctate area of frons almost to base; pronotum with a broad, entire impunctate mid-longitudinal band, the punctural interstices on sides flat, shiny, in parts broader than diameter of punctures; disc of elytra each with a large conspicuous dense fascia of long coppery-red pubescence, and sparser, pale brassy pubescence on humeral angles and lateral parts; large species (18-20 mm). Habitus: fig. 6; male sternite VII unmodified; aedoeagus: figs 6a, 6al ...................................... ignipennis nov.sp.

- Head and pronotum duller, head entirely rugosely punctate; pronotum rugosely punctate, at most with a very narrow, interrupted trace of a mid-longitudinal line.............9

9 Small species (10-12 mm); elytra fuscous, slightly rufescent just behind humeral angles, with sparse long coppery red pubescence covering most of anterior portions of elytra, sometimes extending almost to posterior margins; male sternite VII modified: Habitus: fig.: 7; aedoaegus: figs 7a, 7al ................................................. barbarossa nov.sp.

- Larger species, (14-22 mm)..............................................................................................10

10 Elytra with a bluish tinge, contrasting with black pronotum; terminal abdominal tergites entirely black; length: ca. 17 mm; male sternite VII unmodified. Habitus: fig. 8; aedoeagus: figs 8a, 8al ......................................................................... eleanorae nov.sp.

- Elytra black, concolorous with pronotum, excepting colour of pubescence.....................11

11 Large species, ca. 21 mm; head at widest point (just in front of posterior angles) as wide as elytra (3.8 mm); pronotum with very coarse long vermiculate rugae; elytra depressed, with denser but scattered long pale pubescence laterally, but without distinct fascia of brassy or coppery pubescence; abdominal urites VII and VIII entirely black. Habitus: fig. 9 ............................................................................ aureomontis nov.sp.

- Species either comparably large (19-21 mm), but elytra slightly wider than head, pronotum more homogeneously punctate, without very coarse long vermiculate rugae, and elytra convex, with a conspicuous arcuate line of brassy and coppery pubescence ( N. sarahae nov.sp.), or smaller species (14-17 mm) with elytra very distinctly wider than head...........................................................................................................................12

12 Large species (19-29 mm), head ca. 3.5 mm. Elytra with a shallowly arcuate line of brassy pubescence fringed on inner side with coppery hairs extending from humeral angle, curving inward towards suture and then outwards towards but not reaching postero-lateral angle; sculpture of area between this line and suture obscured by short and dense black pubescence, sculpture laterad of the line coarse, evident and almost glabrous apart from longer sparse black pubescence; posterior 1/3 rd of tergite VII and tergite VIII rufescent; male: sternite VII modified. Habitus: figs 15, 15al............................ .................................................................................................................... sarahae nov.sp.

- Smaller species, 14-17 mm long. Elytra with a diffuse patch of coppery pubescence extending as far as 1/3 rd or ½ the length of elytron, or with scattered silvery pubescence behind humeral angles; sculpture in juxtasutural areas not obscured by dense black pubescence, not differing in appearance from sculpture of lateral parts of disc; tergites VII and VIII entirely black ( N. borneensis , N. calcicola , N. argentifer ) or posterior margin of VII and whole of VIII rufescent........................................................13

13 Elytra with a diffuse patch of coppery pubescence behind humeral angles; pubescence of lateral parts of tergites III-V and VII golden or coppery; male sternite VII modified..14

- Elytra with scattered silvery pubescence; pubescence of lateral parts of tergites III-V and VII silvery; male: sternite VII unmodified; sternite VIII with a very small, shallow emargination; length 16-17 mm. Habitus: fig. 14; aedoeagus: figs14a, 14al ........................ ................................................................................................................. argentifer nov.sp.

14 Smaller species (ca. 14 mm); puncturation of head coarser; male sternite VII (fig. 10c); sternite VIII: fig. 10d; aedeagus: figs 10a, 10al. Habitus: fig. 10........ borneensis CAMERON

- Larger species (16-17 mm); puncturation of head finer; entire dorsal surface of forebody dull; patches of brassy pubescence on anterior parts of elytra monochrome, extending to inner 1/3 rd of elytron; sculpture of abdominal tergites dull, dense, confused............................................................................................................................15

15 Entire dorsal surface duller; brassy pubescence on anterior 2/3rds monochrome, brassy, not mixed with coppery red hairs, often extending to inner half of elytron; sculpture of tergites not shiny-sericeous, more confused......................................................................16

- Entire dorsal surface more shiny; brassy pubescence on anterior 2/3rds of elytra mixed with coppery red hairs, not extending to inner half of elytron; sculpture of tergites shiny-sericeous, with sparse large punctures and clearly defined micro-punctures. Habitus: fig. 13; aedoeagus: figs 13a, 13al.................................................. calcicola nov.sp

16 Puncturation of head, pronotum and lateral of elytra coarser; pronotum longer (2.9) and narrower; patch of brassy pubescence on elytra less extensive; tergite VII shiny, with coarser and sparser punctures. Habitus: fig. 11; aedoeagus: figs11a, 11 al ........................... ................................................................................................................... brendelli nov.sp.

- Puncturation of head, pronotum and lateral half of elytra finer, pronotum shorter (2.7) and less narrowed posteriorly; patch of brassy pubescence more extensive; tergite VII duller, with finer and denser punctures. Habitus: fig. 12; aedoeagus: figs 12a, 12al ............ ................................................................................................................ oxoniensis nov.sp.