Naddia borneensis CAMERON

De, G., 2014, The genus Naddia in Borneo (Coleoptera: Staphylinidae: Staphylininae), Linzer biologische Beiträge 46 (2), pp. 1737-1765 : 1752-1754

Naddia borneensis CAMERON

(Figs 10, 10a, 10al, 10b, 10c)

Naddia borneensis CAMERON 1930: 165 .

Naddia borneensis HAMMOND 1984: 206 .

Naddia borneensis HERMAN 2001: 3353 .

Holotype: (bearing an NHML holotype label: small round white label with a red border): N. BORNEO, BETTOTAN NR. SANDAKAN, Aug. 22 nd 1937; Naddia borneensis CAM. (in NHML).

Additional material: 1 & 1 ex. (abdominal segments 8-10 missing): MALAYSIA, Borneo, Sabah, Ulu Segama Forest Reserve, Danum Valley Conservation Area ca. 10 km from Danum Valley F.C ., Borneo Rainforest Lodge area , Plot 3, 12.III.2004, F.I.Trap, ca. N05˚02 E117˚45/55 Leg. E. Slade & J. Villanueva, OUMNH-2005-062 coll. E. Slade, Oxford University Museum of Natural History (in OUMNH) ; 1: Malaysia, Borneo, Sabah, Ulu Segama Forest Reserve, Yayasan Sabah Logging Concession ca. 10 km from Danum Valley F.C. , Secondary selectively logged forest, 17.IV.2004 F.I.Trap, Coupe 81, Plot 3 ca. N 04˚58.66 E 117˚53.41 Leg. E. Slade & J. Villanueva, OUMNH-2005-062 , coll. E. Slade, Oxford University Museum of Natural History (in OUMNH) ; 1 & 1: MALAYSIA, Sabah, Lahad Datu, Ulu Segama For. Res. Danum Valley F.C. , 04˚57.9’N 117˚48.1’E, 200 m alt., 21.xi.2005, 1˚ Forest, coll. Mann, Slade & Villanueva, Flight Interception Trap, Lowland, Mixed Dipterocarp Forest, West Trail area, OUMNH-2006-051 (in OUMNH, 1 ex. in RCO) ; 1: SABAH, Danum Valley , B.R.L, f.i.t., 14- 16.II.2007, G. de Rougemont (in RCO) ; 1: SABAH, Poring Hot Springs, 500 m, 8.V.1987, Burkhardt–Löbl (in MHNG) ; 7 exx: MALAYSIA, Sabah, Tawau, 8-14.xi.2012, 4.70N 117.52E, 200 m Alt., Coll. C.L. Gray, SAFE project area, F.I.T Riparian forest strip in oil palm, OUMNH-2013-056 (in OUMNH) ; 1: MALAYSIA, Sabah, Tawau, 18-24.x.2012, 4.76N 117.54E, 399 m Alt., Coll. C.L. Gray, SAFE project area, F.I.T. Riparian forest strip in oil palm, OUMNH-2013- 056 (in OUMNH) ; 1: MALAYSIA, Sabah, Tawau, 11-17.x.2012, 4.66N 117.6E, 399 m Alt., Coll. C.L. Gray, SAFE project area, F.I.T Riparian forest strip in oil palm, OUMNH-2013-056 (in LL) .

Redescription: Proportions (of a male from Danum valley): length: 14 mm; length of head: 2.0; total length of head: 2.5; breadth of head: 2.5; length of eye: 0.9; length of temple: 1.1; length of antenna: 3.0; length of pronotum: 2.6; breadth of pronotum: 2.45; length of elytron: 3.0; breadth of elytra: 3.1. Habitus: fig. 10.

Eyes not very prominent because slightly displaced towards dorsal surface of head. Antennomeres 1-3 short, only a little elongate, 4 and 5 sub-globose, 6 to10 increasingly transverse, the penultimate segment as broad as long; segment 11 with conspicuous very dense short white pubescence.

Integument entirely black. Antennae fuscous, legs brownish black. Head and pronotum without coloured pubescence in addition to dark pubescence; lateral setae short; Elytra rather sparsely and irregularly covered in paler hairs mixed with dark pubescence, making them appear brownish; abdominal tergites III-V with elongate lateral fasciae of pale pubescence; tergite VII with two broader but shorter antero-lateral fasciae of silvery pubescence, the fasciae becoming brassy posteriad; tibiae with yellowish pubescence and spines; lateral margins of tarsal segments with reddish pubescence.

Sculpture of head coarser than that of N. brendelli and N. oxoniensis , the interstices forming short irregular longitudinal rugae on disc and frons, the umbilicate punctures still clearly except close to anterior margin; umbilicate punctures of pronotum about the same size as those of head, close, the surface sub-rugose, the interstices tending to form oblique rugae in anterior ¼, the sculpture leaving a very narrow, almost entire vestige of an impunctate mid-longitudinal band; sculpture of elytra confused, consisting of dense umbilicate punctures about as large as those of head and pronotum; puncturation of abdominal tergites dense and fine on middle of tegites III to VI, coarser and sparser on the sides, very sparse on whole of tergite VII, the integuments with dense micro-punctures on anterior half and very faint but discernible transverse micro-striation on posterior half.

Male: sternite VII: (fig. 10c) modified with very long fine pale setae arising from surface in front of sub-circular depression; surface of sternite with fairly dense micro-sculpture, the posterior margin slightly concave; tergite VIII: fig. 10d, the apical margin slightly, very broadly and shallowly emarginate; sternite VIII: fig. 10b, with a small broad emargination, the surface of sternite shiny, with scarcely any microsculpture in anterior half and distinct transverse micro-striation in posterior half; aedoeagus: figs 10a, 10al, the median lobe long in lateral view, curved ventrally, the apex apparently cupuliform; ventral extension of basal sclerite sub-truncate; paramere adpressed, extremely short, the anterior margin with a median notch, in the illustration the paramere is concealed by the large ventrally protruberant basal bulb, but is visible in lateral view.

The type is in poor condition: head and pronotum, and abdomen detached from mesothorax; left protibia and tarsus, right hind leg and segments 5-11 of both antennae missing.

The specimen has been remounted from a pin onto a card and the genital segments and aedoeagus (both in good condition) dissected and glued to the same card.