Orthogeomys Merriam 1895

Wilson, Don E. & Reeder, DeeAnn, 2005, Order Rodentia - Family Geomyidae, Mammal Species of the World: a Taxonomic and Geographic Reference (3 rd Edition), Volume 2, Baltimore: The Johns Hopkins University Press, pp. 859-870 : 864

Orthogeomys Merriam 1895

Orthogeomys Merriam 1895 , N. Amer. Fauna, 8: 23.

Type Species: Geomys scalops Thomas 1894

Species and subspecies: 11 species with 37 subspecies in 3 subgenera:

Subgenus Orthogeomys (Orthogeomys) Merriam 1895

Subgenus Orthogeomys (Heterogeomys) Merriam 1895

Subgenus Orthogeomys (Macrogeomys) Merriam 1895

Species Orthogeomys (Macrogeomys) cavator ( Bangs 1902)

Subspecies Orthogeomys (Macrogeomys) cavator subsp. cavator ( Bangs 1902)

Subspecies Orthogeomys (Macrogeomys) cavator subsp. nigrescens (Goodwin 1943)

Subspecies Orthogeomys (Macrogeomys) cavator subsp. pansa ( Bangs 1902)

Species Orthogeomys (Macrogeomys) cherriei (J. A. Allen 1893)

Subspecies Orthogeomys (Macrogeomys) cherriei subsp. cherriei (J. A. Allen 1893)

Subspecies Orthogeomys (Macrogeomys) cherriei subsp. carlosensis ( Goodwin 1934)

Subspecies Orthogeomys (Macrogeomys) cherriei subsp. costaricensis (Merriam 1895)

Species Orthogeomys (Orthogeomys) cuniculus Elliot 1905

Species Orthogeomys (Macrogeomys) dariensis (Goldman 1912)

Species Orthogeomys (Orthogeomys) grandis ( Thomas 1893)

Subspecies Orthogeomys (Orthogeomys) grandis subsp. grandis ( Thomas 1893)

Subspecies Orthogeomys (Orthogeomys) grandis subsp. alleni Nelson and Goldman 1930

Subspecies Orthogeomys (Orthogeomys) grandis subsp. alvarezi Schaldach 1966

Subspecies Orthogeomys (Orthogeomys) grandis subsp. annexus Nelson and Goldman 1933

Subspecies Orthogeomys (Orthogeomys) grandis subsp. carbo Goodwin 1956

Subspecies Orthogeomys (Orthogeomys) grandis subsp. engelhardi Felten 1957

Subspecies Orthogeomys (Orthogeomys) grandis subsp. felipensis Nelson and Goldman 1930

Subspecies Orthogeomys (Orthogeomys) grandis subsp. guerrerensis Nelson and Goldman 1930

Subspecies Orthogeomys (Orthogeomys) grandis subsp. huixtlae Villa 1944

Subspecies Orthogeomys (Orthogeomys) grandis subsp. latifrons (Merriam 1895)

Subspecies Orthogeomys (Orthogeomys) grandis subsp. nelsoni (Merriam 1895)

Subspecies Orthogeomys (Orthogeomys) grandis subsp. pluto Lawrence 1933

Subspecies Orthogeomys (Orthogeomys) grandis subsp. pygacanthus (Dickey 1928)

Subspecies Orthogeomys (Orthogeomys) grandis subsp. scalops ( Thomas 1894)

Subspecies Orthogeomys (Orthogeomys) grandis subsp. soconuscensis Villa 1949

Subspecies Orthogeomys (Orthogeomys) grandis subsp. vulcani Nelson and Goldman 1931

Species Orthogeomys (Macrogeomys) heterodus ( Peters 1865)

Subspecies Orthogeomys (Macrogeomys) heterodus subsp. heterodus ( Peters 1865)

Subspecies Orthogeomys (Macrogeomys) heterodus subsp. cartagoensis (Goodwin 1943)

Subspecies Orthogeomys (Macrogeomys) heterodus subsp. dolichocephalus (Merriam 1895)

Species Orthogeomys (Heterogeomys) hispidus (Le Conte 1852)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. hispidus (Le Conte 1852)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. cayoensis (Burt 1937)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. chiapensis ( Nelson and Goldman 1929)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. concavus ( Nelson and Goldman 1929)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. hondurensis (Davis 1966)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. isthmicus ( Nelson and Goldman 1929)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. latirostris ( Hall and Alvarez 1961)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. negatus ( Goodwin 1953)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. teapensis (Goldman 1939)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. tehuantepecus (Goldman 1939)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. torridus (Merriam 1895)

Subspecies Orthogeomys (Heterogeomys) hispidus subsp. yucantanensis ( Nelson and Goldman 1929)

Species Orthogeomys (Heterogeomys) lanius ( Elliot 1905)

Species Orthogeomys (Macrogeomys) matagalpae (J. A. Allen 1910)

Species Orthogeomys (Macrogeomys) thaeleri Alberico 1990

Species Orthogeomys (Macrogeomys) underwoodi (Osgood 1931)

Discussion: Russell (1968 a) revised the genus, and included Heterogeomys and Macrogeomys as valid subgenera. Hafner (1991) examined molecular phylogenetics of species in the subgenus Macrogeomys while Sudman and Hafner (1992) examined relationships among species within each of the subgenera. Species limits for those taxa known from single localities or otherwise geogeographically restricted areas are poorly defined.