Janalia jarmilae

Janák, Jiří, 2018, A new genus of the Tympanophorus lineage of the subtribe Anisolinina from Madagascar (Coleoptera: Staphylinidae: Staphylininae: Staphylinini), Zootaxa 4461 (2), pp. 293-299: 295-299

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Janalia jarmilae

sp. nov.

Janalia jarmilae  sp. nov.

( Figs. 1–17View FIGURES 1–6View FIGURES 7–14View FIGURES 15–17)

Type locality. East Madagascar, between Beforona and Ampasimbe, 18° 58' S, 48° 38' E.GoogleMaps 

Type material. Holotype ♂: MADAGASCAR: “E Madagaskar, 2-4 km NNW of RN2 [= route nationale 2 = national road 2], pk 181 [= point kilomètrique = milestone 181], between Beforona and Ampasimbe 500-600 m, 22.-24.2., 18 58 S, 48 38 E, J. Janák lgt. 2004”, “rain forest, forest litter, sifting”, “ HOLOTYPE Janalia jarmilae  gen. nov. sp. nov. J. Janák det. 2018” ( JJRC).

Description. Body length 10.0 mm, forebody length 4.7 mm ( Fig. 1View FIGURES 1–6). Body black, palpi, and antennae dark brown, legs brown, mandibles reddish.

Head ( Fig. 2View FIGURES 1–6) about as long as wide (L/W=1.04), markedly narrowed behind eyes. Temples slightly longer than eyes (R=1.11). Dorsal macropunctures (one side only): two behind antennal base; one at anterior inner border of eye; one medially at midlength of eye; one at posterior inner margin of eye; one lateral (with macroseta) between posterior margin of eye and neck constriction; one medially near neck constriction (with macroseta); three medially between posterior margin of eye and neck constriction. Surface without microsculpture, sparsely and finely punctate and shortly setose. Maxillary palpi with apical palpomere distinctly longer than previous triangular segment, weakly dilated fusiform ( Fig. 6View FIGURES 1–6). Labial palpi with apical palpomere distinctly dilated, not flattened laterally and with broad apical face ( Figs. 4–6View FIGURES 1–6). Ventral surface of head coarsely and widely punctate, with gular setae distant ( Figs. 3, 4View FIGURES 1–6). Antennae short, fifth segment slightly transverse (L/W=0.78), segments 6–9 transverse, tenth segment about two times as wide as long (L/W=0.54). Mandibles relatively short and broad, sharp apically ( Figs. 5, 6View FIGURES 1–6).

Pronotum ( Fig. 2View FIGURES 1–6) slightly longer than wide (L/W=1.06), markedly longer (R=1.40) and wider (R=1.37) than head, with dorsal series consisting of 1+5 fine punctures and lateral series with 1+4 fine punctures, between lateral series and anterior angle with additional fine puncture, surface without microsculpture, sparsely and very finely punctate and shortly setose, punctation finer than that on head, lateral margin with a few relatively short but erect setae. Elytra with dense and coarse punctation, becoming finer at basal depression, shorter than wide (L/W=0.96), longer (R=1.22) and wider (R=1.35) than pronotum. Wings well developed. Scutellum with punctures confluent forming irregular transverse ridges. Abdomen with elongate markedly coarse and moderately dense punctation, all tergites without microsculpture. Legs densely covered with brown setae, all tibiae with lateral spines. Apical metatarsomere longer than previous two tarsomeres.

Male. Sternite 7 ( Fig. 9View FIGURES 7–14) in apical third with depression with modified erect setae ( Figs. 9, 10View FIGURES 7–14). Sternite 8 ( Fig. 9View FIGURES 7–14) with posterior margin slightly emarginate. Sternite 9 bilobed ( Fig. 16View FIGURES 15–17), tergite 10 subtruncate ( Fig. 15View FIGURES 15–17). Aedeagus 0.55 mm long ( Figs. 11–14View FIGURES 7–14); median lobe broad, markedly narrowed apically, paramere short and broad; about 20 sensory peg setae situated confusedly in apical half of paramere ( Fig. 14View FIGURES 7–14).

Female. Unknown.

Differential diagnosis. Janalia jarmilae  gen. nov., sp. nov. is externally similar to Madagascan Trigonopalpus  species and shares with this genus the shape of the apical labial palpomere not laterally flattened, with broad apical face. The new species differs from all these species by long fusiform last maxillary palpomere, distant gular setae and additional generic characters described in the diagnosis of Janalia  gen. nov.

Etymology. The species is dedicated to Jarmila, the wife of my friend and colleague Petr Bulirsch, for her support of mine and Peter’s joint trips to Madagascar and South Africa.

Distribution. Janalia jarmilae  gen. nov. spec. nov. is known only from a small remnant of indigenous forest NNW of National Road between Beforona and Ampasimbe ( Fig. 17View FIGURES 15–17).

Bionomics. The new species was collected by sifting forest litter in a rain forest.