Tympanophorus, Nordmann, 1837

Janák, Jiří, 2018, A new genus of the Tympanophorus lineage of the subtribe Anisolinina from Madagascar (Coleoptera: Staphylinidae: Staphylininae: Staphylinini), Zootaxa 4461 (2), pp. 293-299 : 295

Key to genera of the Tympanophorus lineage

(based on Brunke et al. 2017, modified)

1 Apical labial palpomere slender, fusiform............................................................................................... Tolmerinus Bernhauer, 1923 , Diatrechus Bernhauer, 1911 , Paradiatrechus Cameron, 1944

- Apical labial palpomere distinctly dilated apically and almost always securiform ( Figs. 4–6 and Figs. 4E, F in Brunke et al. 2017)............................................................................................... 2

2 Apical labial palpomere laterally flattened with narrow apical face ( Fig. 4E in Brunke et al. 2017)..................... 3

- Apical labial palpomere not laterally flattened, with broad apical face ( Figs. 4–6 )................................... 4

3 Labrum expanded, lobes less than twice as wide as long ( Fig. 4E in Brunke et al. 2017); apical maxillary palpomere weakly to moderately expanded subapically or apically, apex never pointed ( Fig. 4B, E in Brunke et al. 2017); pronotum punctate across entire disc.................................................................... Barygnathus Bernhauer, 1902

- Labrum not expanded, lobes distinctly more than twice as wide as long (as in Figs. 2C, 4F in Brunke et al. 2017); apical maxillary palpomere evenly converging posteriad to pointed apex ( Fig. 4A in Brunke et al. 2017); pronotum with median impunctate line......................................................................... Amelinus Bernhauer, 1915

4 Apical maxillary palpomere fusiform, not conspicuously constricted apically, at least twice as long as previous segment ( Fig. 4C in Brunke et al. 2017, Fig. 6 ).......................................................................... 5

- Apical maxillary palpomere conspicuously constricted apically, at most slightly longer than previous segment ( Figs. 4D, F in Brunke et al. 2017).................................................................................... 7

5 Head with temples nearly straight and strongly converging to neck; body with areas of long golden pubescence (Fig. 32 in Schillhammer, 2002); male unknown.................................................. Bombylodes Fauvel, 1904

- Head with temples rounded or only slightly converging to neck ( Fig. 3B in Brunke et al. 2017, Figs. 1–3 ); body without conspicuous areas of golden pubescence....................................................................... 6

6 Paramere with long apex and distinct expansion at midlength ( Figs. 1 –19 in Schillhammer 2002)... Pammegus Fauvel, 1895

- Paramere short and broad ( Figs. 11–14 )...................................................... Janalia gen. nov.

7 Ventral surface of head with gular setae approximate ( Fig. 4G in Brunke et al. 2017); prosternum not projected ventrad.................................................................................. Trigonopalpus Cameron, 1951

- Ventral surface of head with gular setae distant ( Fig. 4H in Brunke et al. 2017); pronotum projected ventrad......................................................................................... Tympanophorus Nordmann, 1837

Figures

FIGURES 1–6. Janalia jarmilae gen. nov., sp. nov. 1, habitus; 2, forebody; 3, head ventral; 4, head ventro-lateral; 5, mouthparts dorsal; 6, mouthparts ventral.

FIGURES 7–14. Janalia jarmilae gen. nov., sp. nov. 7, 8, mesoventrite; 9, sternites 6-8; 10, abdominal segments 6-8 lateral; 11, aedeagus lateral; 12, aedeagus ventro-lateral; 13, aedeagus ventral; 14, aedeagus ventral in transmitted light. 11–14, same scale bars.