Rhagovelia angustipes, Uhler, 1894
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publication ID |
https://doi.org/ 10.11646/zootaxa.5361.3.4 |
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publication LSID |
lsid:zoobank.org:pub:89A338C1-1DD5-439E-9CBE-74151FC5A4B5 |
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DOI |
https://doi.org/10.5281/zenodo.10167910 |
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persistent identifier |
https://treatment.plazi.org/id/587AC751-B57B-A200-FF42-F895BCACFB33 |
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treatment provided by |
Plazi |
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scientific name |
Rhagovelia angustipes |
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Key to the species of the Rhagovelia angustipes View in CoL complex from Colombia
[Modified from Galindo-Malagón et al. (2021); apterous males and/or females are required or recommended for several of the steps; species of the bisignata and hambletoni groups are treated together because macropterous specimens necessary to distinguish them can be rare; includes R. medinae Galindo-Malagón , Mondragón-F., Morales & Moreira, 2022, whose apterous pronotum does not fit perfectly in the definition of the complex but that is assigned to the hambletoni group; also note that R. guachiconoense Padilla-Gil, 2019 might be a synonym of R. longipes Gould, 1931 ]
1 Monomorphic apterous; middle tarsus biarticulated; found mainly on brackish or salt water (e.g., mangroves, estuaries)............................................................................................ [ salina View in CoL group] 2
- Dimorphic apterous or macropterous; middle tarsus triarticulated; found mainly on freshwater (e.g., streams, rivers)............................................................................... [ bisignata View in CoL and hambletoni groups] 7
2 Eye covered by abundant setae........................................................................... 3
- Eye with one or two setae only........................................................................... 4
3 Body length 2.20–2.40 mm in the male and 3.30–3.50 mm in the female; posterior surface of male hind femur with a row of 3–7 spines; male paramere as in Fig. 11D View FIGURE 11 ......................................... Rhagovelia plumbea Uhler, 1894 View in CoL
- Body length ~ 1.75 mm in the male and ~ 2.90 mm in the female; posterior surface of male hind femur with a row of 6–10 spines; male paramere as in Fig. 11E View FIGURE 11 ................................. Rhagovelia tintipan Molano, Morales & Moreira, 2018 View in CoL
4 Male fore tibia straight or slightly curved ( Fig. 13B View FIGURE 13 ); male abdominal segment VIII large, longer dorsally than abdominal mediotergite VII ( Fig. 5B View FIGURE 5 ); male paramere strongly sculptured ( Fig. 11B View FIGURE 11 ); male proctiger as in Fig. 12B View FIGURE 12 ; dorsum of female abdominal terga VI –VIII (sometimes also V) shiny black centrally...................................................................................................... Rhagovelia colombiana (J. Polhemus & Manzano, 1992) View in CoL
- Male fore tibia strongly curved ( Fig. 13A View FIGURE 13 ); male abdominal segment VIII small, approximately equal in length dorsally to abdominal mediotergite VII (as in Fig. 5A View FIGURE 5 ); male paramere shape simpler ( Figs. 11A, 11C, 11E View FIGURE 11 ); male proctiger not as above; at most dorsum of female abdominal terga VII–VIII shiny black or brown centrally (as in Fig. 5C View FIGURE 5 )..................... 5
5 Body length ~3.00 mm in the male and ~ 4.20 mm in the female; male paramere and proctiger as in Figs. 11A View FIGURE 11 , 12A View FIGURE 12 ................................................................... Rhagovelia arcuata (J. Polhemus & Manzano, 1992) View in CoL
- Body length 2.20–2.60 mm in the male and 3.00– 3.20 mm in the female; male paramere and proctiger not as above....... 6
6 Posterior surface of male hind femur with about 6 spines beyond middle; male abdominal sterna without median carina; female abdominal laterotergites slightly elevated ( Fig. 5E View FIGURE 5 )............................ Rhagovelia rosarensis Padilla-Gil, 2010 View in CoL
- Posterior surface of male hind femur with about 5 spines basally, followed by one long spine before middle, then about 13 smaller spines toward apex; male abdominal sterna VII–VIII with weak median carina; female abdominal laterotergites vertical or nearly so ( Fig. 5D View FIGURE 5 )......................................... Rhagovelia nuqui Molano, Morales & Moreira, 2018 View in CoL
7 Hind femur without spines in both sexes ( Fig. 13D View FIGURE 13 ).......................................................... 8
- Hind femur with spines in both sexes ( Figs. 13E, 13F View FIGURE 13 )....................................................... 10
8 Dorsum of abdominal terga V–VIII shiny black centrally; male paramere and proctiger as in Figs. 11V View FIGURE 11 , 12V View FIGURE 12 ............................................................................. Rhagovelia guachiconoense Padilla-Gil, 2019
- At most dorsum of abdominal segments VII–VIII shiny black centrally; male paramere and proctiger not as above........ 9
9 Antennomere II shorter than III; fore and hind coxae and trochanters dark-brown to black, rarely brown to yellow; male fore tibia very thin for about 3/4 of length, then expanding toward apex ( Fig. 13C View FIGURE 13 ); paramere and proctiger as in Figs. 11O View FIGURE 11 , 12O View FIGURE 12 ........................................................................... Rhagovelia longipes Gould, 1931 View in CoL
- Antennomere II longer than III; fore and hind coxae and trochanters yellow ( Fig. 7A View FIGURE 7 ); male fore tibia thicker and more uniform throughout length; paramere and proctiger not as above.......................... Rhagovelia caunapi Padilla-Gil, 2015
10 Male metasternum and abdominal sterna II–IV strongly swollen, with brushes of long golden setae medially ( Fig. 7B View FIGURE 7 ); lateral margins of male abdominal segment VIII clearly divergent posteriorly ( Fig. 7B View FIGURE 7 ); male paramere and proctiger as in Figs. 11L View FIGURE 11 , 12L View FIGURE 12 . .............................................................. Rhagovelia gastrotricha Padilla-Gil, 2011 View in CoL
- Male metasternum and abdominal sterna II–IV not strongly swollen, covered only by short setae in most species; male abdominal segment VIII subcylindrical, with lateral margins parallel or bowed, but not clearly divergent posteriorly; male paramere and proctiger not as above................................................................................. 11
11 Male hind trochanter with spines; male hind femur much thicker than middle femur, enormously incrassate in most specimens ( Figs. 6E View FIGURE 6 , 13F View FIGURE 13 ); male paramere and proctiger as in Figs. 11I View FIGURE 11 , 12I View FIGURE 12 ................ Rhagovelia calopa Drake & Harris, 1927 View in CoL
- Male hind trochanter without spines; male hind femur at most slightly thicker than middle femur, never enormously incrassate (as in Figs. 13D, 13E View FIGURE 13 ); male paramere and proctiger not as above.............................................. 12
12 Apterous pronotum uniquely conformed, longer than dorsal eye length, partially covering mesonotum, with posterior margin slightly convex and a transverse sulcus on the posterior region, giving the impression of a double posterior margin ( Fig. 7E View FIGURE 7 ); male abdominal sternum VII with pair of projections bearing tufts of brown setae ( Fig. 7F View FIGURE 7 ); male paramere and proctiger as in Figs. 11W View FIGURE 11 , 12W View FIGURE 12 . ..................... Rhagovelia medinae Galindo-Malagón , Mondragón-F., Morales & Moreira, 2022
- Apterous pronotum typically conformed, shorter than dorsal eye length, with posterior margin straight or slightly concave (as in Figs. 1A, 1C View FIGURE 1 ); male abdominal sternum VII without pair of projections bearing tufts of brown setae; male paramere and proctiger not as above................................................................................. 13
13 Male abdominal sternum VI or VII with a projection or spine................................................. 14
- Male abdominal sterna VI and VII lacking projections and spines.............................................. 15
14 Male abdominal sternum VI with a small, triangular projection on posterior margin ( Fig. 1E View FIGURE 1 ); male abdominal sternum VII lacking a spine; male paramere and proctiger as in Figs. 2C, 2D View FIGURE 2 ; female abdominal laterotergites folded over mediotergites, strongly convergent, almost touching over between segments V and VI, then divergent toward apex of body ( Figs. 1C, 1D View FIGURE 1 ).............................................................................. Rhagovelia andradei sp. nov.
- Male abdominal sternum VI lacking a posterior projection; male abdominal sternum VII with a stout, slightly curved spine; male paramere and proctiger as in Figs. 11S View FIGURE 11 , 12S View FIGURE 12 ; female abdominal laterotergites vertical anteriorly, slightly folded over mediotergites for posterior most three segments.................................... Rhagovelia spinosa Gould, 1931 View in CoL
15 Body length 2.60–2.90 in the male and 2.80–3.10 in the female................................................ 16
- Body length 3.00– 4.20 in the male and 3.20–4.70 in the female................................................ 20
16 Male hind tibia lacking spines and apical spur (apex of tibia can sometimes bear a tuft of setae, but never a spur)........ 17
- Male hind tibia with short spines and an apical spur (as in Fig. 2A View FIGURE 2 )............................................. 18
17 Lateral margins of male abdomen more evenly tapering to apex ( Fig. 8E View FIGURE 8 ); male paramere and proctiger as in Figs. 11T View FIGURE 11 , 12T View FIGURE 12 ; female hind femur thinner ( Fig. 10A View FIGURE 10 ); female abdomen elongated ( Fig. 10A View FIGURE 10 )...... Rhagovelia tantilla Drake & Harris, 1933 View in CoL
- Lateral margins of male abdomen more strongly bowed on segments IV–V ( Fig. 8A View FIGURE 8 ); male paramere and proctiger as in Figs. 11P View FIGURE 11 , 12P View FIGURE 12 ; female hind femur thicker ( Fig. 9D View FIGURE 9 ); female abdomen short and robust ( Fig. 9D View FIGURE 9 )............................................................................ Rhagovelia molanoi Galindo-Malagón, Morales & Moreira, 2021
18 All coxae and trochanters yellowish ( Fig. 7C View FIGURE 7 ); male abdominal sterna II–V with median comb of long golden setae ( Fig. 7C View FIGURE 7 ); male abdominal segment VIII large and robust ( Fig. 7C View FIGURE 7 ); male paramere and proctiger as in Figs. 11N View FIGURE 11 , 12N View FIGURE 12 ; lateral margins of female abdomen sinuous ( Fig. 9C View FIGURE 9 ); posterior margin of female abdominal tergum VIII with long setae................................................................... Rhagovelia graziae Galindo-Malagón, Morales & Moreira, 2021
- Middle trochanter dark-brown to black; male abdominal sterna only with short setae; male abdominal segment VIII small (as in Fig. 8C View FIGURE 8 ); male paramere and proctiger not as above; lateral margins of female abdomen more evenly tapering or bowed (as in Fig. 9E View FIGURE 9 ); posterior margin of female abdominal tergum VIII without long setae................................. 19
19 Male hind femur with 9–11 spines; male hind tibia with 11 short spines and an apical spur; lateral margins of male abdomen more evenly tapering to apex ( Figs. 8C View FIGURE 8 ); male paramere and proctiger as in Figs. 11Q View FIGURE 11 , 12Q View FIGURE 12 ; female hind femur with 6–7 spines; female abdomen more elongated, with laterotergites vertical or nearly so ( Fig. 9E View FIGURE 9 )..... Rhagovelia rosensis Padilla-Gil, 2011 View in CoL
- Male hind femur with 4–7 spines; male hind tibia with 2 short spines and an apical spur; lateral margins of male abdomen more strongly bowed on segments IV–V ( Fig. 8D View FIGURE 8 ); male paramere and proctiger as in Figs. 11R View FIGURE 11 , 12R View FIGURE 12 ; female hind femur with 4–5 spines; female abdomen more robust, with laterotergites slightly elevated ( Fig. 9F View FIGURE 9 )...................................................................................................... Rhagovelia santanderi Padilla-Gil, 2015
20 Body length 3.00– 3.40 in the male and 3.20–3.75 in the female................................................ 21
- Body length 3.50–4.20 in the male and 4.00– 4.70 in the female................................................ 25
21 Male hind tibia only with an apical spur, without small spines throughout length; male paramere and proctiger as in Figs. 11F View FIGURE 11 , 12F View FIGURE 12 . .................................................................... Rhagovelia angustipes Uhler, 1894 View in CoL
- Male hind tibia with small spines throughout length and an apical spur; male paramere and proctiger not as above....... 22
22 Male hind femur thicker, shorter than hind tibia (ratio ~0.90/1.00) ( Fig. 8B View FIGURE 8 ); male hind femur with 4–5 spines; only abdominal tergum VIII shiny black..................................................... Rhagovelia penta Padilla-Gil, 2015
- Male hind femur thinner, longer than hind tibia (ratio ~1.05/1.00) (as in Fig. 6A View FIGURE 6 ); male hind femur with at least 6 spines; central shiny black areas at least on dorsum of abdominal segments VII–VIII........................................... 23
23 Male hind femur slightly surpassing apex of abdomen, with 6 spines ( Fig. 6D View FIGURE 6 ); male paramere and proctiger as in Figs. 11H View FIGURE 11 , 12H View FIGURE 12 ; female hind femur with 4 spines........................................ Rhagovelia calceola Padilla-Gil, 2011 View in CoL
- Male hind femur distinctly surpassing apex of abdomen, with at least 8 spines (as in Fig. 8F View FIGURE 8 ); male paramere and proctiger not as above; female hind femur with at least 7 spines.......................................................... 24
24 Middle coxa dark-brown to black; male hind femur with at least 10 spines; male paramere and proctiger as in Figs. 11U View FIGURE 11 , 12U View FIGURE 12 ; female abdominal mediotergites approximately in the same horizontal plane ( Fig. 10B View FIGURE 10 ); female abdominal laterotergites horizontal or slightly elevated ( Fig. 10B View FIGURE 10 )...................................... Rhagovelia tenuipes Champion, 1898 View in CoL
- Middle coxa yellow (as in Fig. 6B View FIGURE 6 ); male hind femur with 8 spines; male paramere and proctiger not as above; female abdominal mediotergites I–III raised, IV–VI depressed, VII horizontal, tergum VIII declining posteriorly ( Fig. 9A View FIGURE 9 ); female abdominal laterotergites elevated, more strongly on last two segments ( Fig. 9A View FIGURE 9 )........... Rhagovelia barbacoensis Padilla-Gil, 2015
25 Fore and hind trochanters, and middle coxa yellow ( Fig. 10C View FIGURE 10 ); male hind femur shorter and thicker, with 6–7 spines ( Fig. 6F View FIGURE 6 ); male paramere and proctiger as in Figs. 11M View FIGURE 11 , 12M View FIGURE 12 . ............................. Rhagovelia grandis Padilla-Gil, 2011 View in CoL
- Fore and hind trochanters, and middle coxa black; male hind femur longer and thinner, with at least 10 spines (as in Fig. 6C View FIGURE 6 ); male paramere and proctiger not as above................................................................. 26
26 Legs with abundant long black setae; male paramere and proctiger as in Figs. 11G View FIGURE 11 , 12G View FIGURE 12 ; female abdominal laterotergites strongly elevated, almost touching at apex of last segment, which bears abundant brown setae posteriorly ( Fig. 9B View FIGURE 9 )..................................................... Rhagovelia boyacensis Galindo-Malagón, Morales & Moreira, 2021
- Legs without abundant long black setae; male paramere and proctiger not as above; female abdominal laterotergites not as above.............................................................................................. 27
27 Central shiny black areas on dorsum of abdominal segments VII–VIII in the male and VI–VIII in the female; male hind femur with 14–16 spines; male paramere and proctiger as Figs. 11J View FIGURE 11 , 12J View FIGURE 12 ................... Rhagovelia cardia Padilla-Gil, 2011 View in CoL
- Central shiny black areas on dorsum of abdominal segments V–VIII in both sexes; male hind femur with 10 spines; male paramere and proctiger as in Figs. 11K View FIGURE 11 , 12K View FIGURE 12 ................................. Rhagovelia cimarrona Padilla-Gil, 2011 View in CoL
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Mykotektet, National Veterinary Institute |
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Royal British Columbia Museum - Herbarium |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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