Orchestina clavigera, Henrard & Jocqué, 2012

Henrard, Arnaud & Jocqué, Rudy, 2012, 3284, Zootaxa 3284, pp. 1-104 : 24-28

Orchestina clavigera new species

Figures 57–88, 606

TYPE: Male holotype: Kenya, Kakamega Forest , 0.21666°, 34.90000°, advanced secondary forest, canopy of Heinsenia diervilleoides, Jan. 01- Feb. 01, 2002, Freund W. ( ZFMK PBI _OON 33552) .

ETYMOLOGY: The specific name refers to the club-shaped (Latin clava) first tibia in the male.

DIAGNOSIS: Both males and females have a distinctive color pattern of the labium, dark with two pale spots ( Figs. 58, 65), the sternum marginally with a large dark band ( Fig. 58, 65) and a remarkably globular abdomen. Males of O. clavigera are easily recognized by the club-shaped anterior tibia ( Fig. 59). Females are characterized by the tube shaped AUS in the genitalia ( Figs. 69, 70).

These characters are shared with O. microfoliata n. sp. from which the females cannot be separated. However, females of O. clavigera n. sp. and O. microfoliata can easily be distinguished from those of O. gibbotibialis n. sp. in which the sternum has dark stripes radiating from the centre in the anterior half of sternum.

MALE (PBI_OON 8910). Total length 1.10. CEPHALOTHORAX: Carapace ( Figs. 57, 58, 60, 62) brown, broadly oval in dorsal view (CW/CL ≈ 0.83), pars cephalica slightly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width. Clypeus vertical, slightly concave in lateral view. Eyes ALE-PME touching, PME circular; PLE-PME separated by less than PME radius. Sternum ( Figs. 58, 74) as long as wide, yellow-brown, anterior margin unmodified, posterior margin extending posteriorly beyond anterior edges of coxae IV as single extension, with tiny dark spots along margin forming a large darker band, anterior margin slightly concave with transverse depression at base of labium; setae abundant, evenly scattered. Mouthparts ( Figs. 58, 75): Chelicerae, endites and labium pale orange. Chelicerae ( Figs 62) slightly divergent. Labium ( Figs. 74–78) triangular, anterior margin indented at middle, sub distal region with gutter-shaped depression, same as sternum in sclerotization, posterior part with tiny sepia spots delimiting two adjacent, oval areas; with 6 or more setae on anterior margin, subdistal region provided anteriorly with group of five needle like setae, medially with one pair of thin, symmetric, flattened setae in centre, followed posteriorly by one pair of thin asymmetric flattened setae, in a slightly oblique longitudinal row. Endites ( Figs. 75, 76) serrula absent, anteromedian tip unmodified, same as sternum in clerotization. ABDOMEN ( Figs. 57, 60): globular; dorsum soft portions pale white, without color pattern. Pedicel unmodified, scutum absent, but abdomen not extending anterior of pedicel. LEGS: yellow; patella plus tibia I as long as carapace (TL/CL ≈ 1,15), proximal part of tibia I ( Figs. 59, 61, 79 –82) strongly swollen with dorsal part slightly depressed and provided with oblique, longitudinal row of pores, visible with SEM. Tarsus I superior claws ( Figs. 83, 84) with eight teeth on lateral surface of proclaw, six teeth on median surface of proclaw, eight teeth on lateral surface of retroclaw, six teeth on median surface of retroclaw. Tarsal organ I pear shaped, ( Fig. 85). GENITALIA: Palp ( Figs. 61, 66–68) proximal segments white; patella attached to tibia sub-basally; tibia almost cylindrical, slightly more than two times as wide as femur; cymbium yellow-brown, without distal patch of setae; bulb white, stout, tapering apically; embolus dark, slender, needle shaped, as long as bulbus, turned inward, forming almost a right angle with bulbus axis.

FEMALE (PBI_OON 33851). As in male except as noted. Total length 1.48. CEPHALOTHORAX: Carapace ( Figs. 63–64) broadly oval in dorsal view (CW/CL ≈ 0.78); Clypeus sloping forward in lateral view. Mouthparts ( Fig. 65): Chelicerae, endites and labium yellow-brown. Endites serrula present in single row. ABDOMEN (63– 65): without long posterior extension, abdomen extending anterior of pedicel; dorsum soft portions with dark posterior end. Spinnerets ( Fig. 26–28): ALS with five spigots, PMS with two spigots, PLS with three spigots. LEGS: proximal part of tibia I not modified. GENITALIA ( Figs. 69–73, 86–88): with straight tubular structure (AUS), visible through cuticle; in ventral view strongly sclerotized and darker anteriorly; in lateral view clearly folded back over almost half its length; in lateral view, dorsal and ventral parts separated in anterior half, adjacent in posterior half, anterior quarter medially provided with small, faintly sclerotized transverse band (Pr). In dorsal view, lateral extensions (Ex) distally strongly widened and provided with a deep indentation forming two prongs; median one indented at extremity; AUS tubular, consisting of dorsal and ventral parts, surpassing extensions by twice their length, curved over 180°; its ventral part provided with short transverse lateral protrusions situated at 3/4 of the total length; dorsal and ventral parts separated in anterior half, adjacent in posterior half. ARe probably fused with AUS anteriorlyAnterior receptaculum not visible in our preparations; posterior receptaculum absent.

MATERIAL EXAMINED: KENYA: Kakamega Forest , canopy, 0.21666°, 34.90000°, Feb. 07, 1999 to Feb. 11, 1999, Wagner T., 4♂ paratypes ( MRAC 228851 PBI _ OON 8905 ); as previous, 3♀ paratypes ( MRAC 228870 PBI _OON 33630); as previous, 1♀ paratype ( MRAC 228856 PBI _OON 33851); as previous, 1♀ paratype ( ZFMK PBI _OON 33533) ; as previous, 1♀ paratype ( ZFMK PBI _OON 33534) ; as previous, 2♂ paratypes ( MRAC 228854 PBI _ OON 8906 ) ; as previous, 1♂ paratype ( MRAC 228848 PBI _ OON 8907 ) ; as previous, 1♂ paratype ( MRAC 228852 PBI _ OON 8908 ) ; as previous, 1♂ paratype ( MRAC 228845 PBI _ OON 8909 ) ; as previous, 1♂ paratype ( MRAC PBI _ OON 8910 ) ; as previous, 1♀ paratype ( MRAC PBI _OON 33898) ; as previous, 1♀ paratype ( MRAC PBI _OON 33148) ; as previous, Feb. 07, 1999 to Feb. 11, 1999, Freund W., 3♀ paratypes ( ZFMK PBI _OON 33553) ; as previous, 3♂ paratypes ( ZFMK PBI _OON 33556) ; as previous, middle-aged secondary forest, canopy, Jan. 02, 2002, Freund W., 3♂ ( MRAC 228792 PBI _OON 16786) ; as previous, Feb. 01, 2002, 1♂ paratype ( ZFMK PBI _OON 33370) ; as previous, Sept. 01, 2001, 1♀ paratype ( MRAC PBI _OON 33432) ; as previous, 2♂ paratypes ( ZFMK PBI _OON 33371) ; as previous, Feb. 01, 2003, 1♂ paratype ( ZFMK PBI _OON 33372) ; as previous, Oct. 09, 2001, 1♂ paratype ( ZFMK PBI _OON 33373) ; as previous, Feb. 01, 2002 ,, 1♂ paratype ( ZFMK PBI _OON 33374) ; as previous, Oct. 09, 2002, 2♀ paratypes ( MRAC 228793 PBI _OON 16787) ; as previous, Jan. 01, 2003 to Feb. 01, 2003, 3♂ paratypes ( ZFMK PBI _OON 33363) ; as previous, 1♀ paratype ( ZFMK PBI _OON 33364) ; as previous, Feb. 07, 1999 to Feb. 11, 1999, 4♂ paratypes ( MRAC 228846 PBI _OON 33415) ; as previous, canopy, Feb. 07, 1999 to Feb. 11, 1999, 1♀ paratype ( ZFMK PBI _OON 33366) ; as previous, 1♀ paratype ( ZFMK PBI _OON 33367) ; as previous, Jan. 01, 2002 to Feb. 01, 2002, 1♀ paratype ( ZFMK PBI _OON 33531) ; as previous, Oct. 09, 2001, 1♀ paratype ( ZFMK PBI _OON 33532) ; as previous, 2♀ paratypes ( ZFMK PBI _OON 33368) ; as previous, Wagner T., 3♀ paratypes ( ZFMK PBI _OON 33369) ; as previous, 1♂ paratype ( ZFMK PBI _OON 33375) ; as previous, 2♂ paratypes ( ZFMK PBI _OON 33376) ; as previous, 1♂ paratype ( MRAC 228870 PBI _OON 33408) ; Jan. 01, 2003 to Feb. 01, 2003, 2♂ paratypes ( ZFMK PBI _OON 33546) ; as previous 1♂ paratype ( ZFMK PBI _OON 33557) ; as previous, 2♂ paratypes ( ZFMK PBI _OON 33554) ; as previous, 1♂ paratype ( ZFMK PBI _OON 33547) ; as previous, 2♂ paratypes ( ZFMK PBI _OON 33548) ; as previous, Sept. 01, 2001 to Oct. 01, 2001, 1♂ paratype ( ZFMK PBI _OON 33551) ; as previous, Jan. 01, 2002, 1♂ paratype ( ZFMK PBI _OON 33558) ; as previous, Feb. 01, 2002, 1♀ paratype ( MRAC 228850 PBI _OON 33849) ; as previous, 1♀ paratype ( MRAC 228855 PBI _OON 33850) ; as previous, advanced secondary forest, canopy, Feb. 01, 2003, Freund W., 1♂ paratype ( MRAC 228849 PBI _OON 33414) ; as previous, Jan. 01, 2003 to Feb. 01, 2003, 4♀ paratypes ( ZFMK PBI _OON 33545) ; as previous, 2♂ paratypes ( ZFMK PBI _OON 33555) ; as previous, 1♂ ( ZFMK PBI _OON 33574) ; as previous, old secondary forest, canopy, 1♂ paratype ( MRAC 228844 PBI _OON 33416) .

DISTRIBUTION: Known only from the Kakamega Forest in western Kenya ( Fig. 606).