Phanaeus (Notiophanaeus) pyrois Bates, 1887

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 214-217

Phanaeus (Notiophanaeus) pyrois Bates, 1887 Plate 47C

Phanaeus pyrois Bates, 1887: 58 (original description. Type locality: Nicaragua, Chontales; Costa Rica: Panama, Bugaba, Chiriqui Volcano, 2000-3000 feet [= 610-915 m]; South America, Colombia).

Phanaeus pyrois : Nevinson 1892: 6 (catalog, distribution); Gillet 1911a: 85 (complete list of species); d’ Olsoufieff 1924: 37 (characters in key); 93 (comment, distribution), 152 (distribution); Blackwelder 1944: 210 (list of species from Latin America); Medina et al. 2001: 140 (cited for Colombia); Krajcik 2012: 204 (complete list of species); Moctezuma and Halffter 2017: 55 (characters in key, cited for Ecuador); Kohlmann et al. 2018: 79 (figures 8b and d), 83 (characters in key, cited for Ecuador).

Phanaeus (Notiophanaeus) pyrois : Edmonds 1994: 41 (characters in key), 45 (redescription), 103 (distribution); Vitolo 2000: 597 (characters in key); Vitolo 2004: 284 (diagnosis); Carvajal et al. 2011: 322-323 (cited for Ecuador); Edmonds and Zídek 2012: 13 (characters in key), 53 (distribution figure 143); Bezdek and Hajek 2013: 432 (cited as junior subjective synonym of Phanaeus (Notiophanaeus) pyrois Bates, 1887. Catalog of the types of the NMPC); Chamorro et al. 2018: 84 (figure 7D), 98 (cited for Ecuador); Arnaud 2018: 4 (comment), 5 (figures 2e-d).

Phanaeus blanchardi d’Olsoufieff, 1924: 37 (characters in key), 92 (original description, cited for Ecuador); Vulcano and Pereira 1967: 575 (characters in key); Martínez and Pereira 1967: 68 (synonym of Phanaeus (Phanaeus) funereus Balthasar, 1939. Cited for Ecuador); Edmonds, 1994: 45 (synonym of Phanaeus pyrois Bates, 1887); Vitolo 2004: 284 (cited as synonym for Phanaeus pyrois Bates, 1887); Carvajal et al. 2011: 320-321 (cited for Ecuador); Edmonds and Zídek, 2012: 5 (cited as jr. prim. hom. of blanchardi Harold. Permanently unavailable, valid name pyrois Bates), 13 (comment).

Phanaeus (s. str.) funereus Balthasar, 1939: 241 (original description, cited for Ecuador).

Phanaeus funereus : Blackwelder 1944: 209 (list of species from Latin America); Martínez and Pereira 1967: 68 (distribution, comment); Edmonds 1994: 45 (synonym of Phanaeus pyrois Bates, 1887); 103 (distribution); Edmonds and Zídek 2012: 5 (cited as junior synonym for Phanaeus pyrois Bates, 1887).

Phanaeus (Notiophanaeus) pyrois funereus : Arnaud 2002a: 97 (diagnosis); Carvajal et al. 2011: 322-323 (cited for Ecuador).

Type specimens.

Phanaeus pyrois Bates, 1887. The lectotype (♂) is deposited at the NHML (ex coll. H. Bates). Locality: Chontales, Nicaragua, examined.

Lectotype (♂): "Chontales / Nicaragua / T. Belt. [p]", "P. pyrois ♂ [hw]", "B.C.A. / p. 58, sp 8. [p]", "Sp. figured. [p]", "Phanaeus / pyrois. Bates / Lectotype ♂ / P. ARNAUD DET 1980 [hw and p, red margin]".

Phanaeus blanchardi d’Olsoufieff, 1924. The lectotype (♂) is deposited at the MNHN (see Arnaud 1982a: 116). Locality: Colombie Vallée de Cauca [= Colombia, Valle del Cauca], not examined.

Phanaeus (s. str.) funereus Balthasar, 1939. The holotype (♂) is deposited at the NMPC (ex coll. V. Balthasar). Locality: Pucay [= Bucay], examined.

Holotype (♂): "ECUADOR / Pucay / F.O., 10.6.05 [p]", "♂ [hw]", "H. Blut Determ. / Phanaeus / blanchardi Olsuf. [hw and p]", "Felsche det. / Phanaeus / pyrois Bts. [hw and p]", "Ph. funereus / m. n.sp. / Dr. v. Balthasar det. [hw and p]", "Typus [p, red label, black margin]", "Mus. Nat. Pragae / Inv. [p], 26347 [hw, orange label]", "ex coll. V. Balthasar / National Museum Prague / Czech Republic [p]".

Distribution.

Colombia and Ecuador.

Records examined.

BOLIVAR: Altamira de Echeandia, 520-720 m (1 specimen MUTPL). CARCHI: Tobar Donoso, 300 m (5 specimens MECN). ESMERALDAS: Colón del Onzole (1 specimen MQCAZ; 8 specimens MECN); Chisprero (5 specimens MECN); E C Río Canandé, 390 m (2 specimens MQCAZ); Jeyambi PMFC (1 specimen MUTPL; 4 specimens MECN); Kumanii Lodge, 40 m (1 specimen MQCAZ); Majua (1 specimen CEMT; 7 specimens MECN); Gualpi El Pajonal (7 specimens MECN); Palma Real (1 specimen CEMT; 5 specimens MECN); Playa de Oro Estero Pote, 200 m (6 specimens CEMT; 11 specimens MECN); Playa de Oro La Tabla (7 specimens MECN; 2 specimens MQCAZ); Playa de Oro Playa Rica (4 specimens MECN); Playa de Oro Padre Santo (1 specimen MQCAZ; 9 specimens MECN); Salto del Bravo (5 specimens MECN); Tjespi (7 specimens MECN); Tjespi río Zapallo (1 specimen MQCAZ). GUAYAS: Guayaquil, 50 m (1 specimen MQCAZ); Pucay [= Bucay] (1 specimen NMPC). IMBABURA: El Chontal, El Cauchero, 900 m (1 specimen MUTPL); Lita, 680 (3 specimens MECN). LOS RÍOS: Quevedo Pichilingue (1 specimen CEMT). MANABÍ: Puerto López, Guale, 310 m (1 specimen MUTPL); Puerto López, Las Tunas, 200 m (1 specimen MUTPL); Puerto López, Puerto Rico, 110 m (1 specimen MUTPL). PICHINCHA: Guayabilla, 515 m, Río Guayllabamba, Manduriacus (1 specimen CEMT); El Encuentro, 620 m, San Miguel de los Bancos (3 specimens MUTPL); El Tigre Río Guayllabamba, Pedro Vicente Maldonado (2 specimens MUTPL); Llurimaguas, 290 m, Río Guayllabamba, Pedro Vicente Maldonado (1 specimen MUTPL); Mangaloma, 820 m, San Miguel de los Bancos (1 specimen MUTPL); Mindo, 1500 m (2 specimens MECN); San Roque, 580 m, Río Guayllabamba, Pedro Vicente Maldonado (2 specimens MUTPL); Tortugo Río Guayllabamba, 450 m, Pedro Vicente Maldonado (1 specimen MUTPL). SANTA ELENA: La Rinconada, 10 m (1 specimen MQCAZ); Olón 50 m (2 specimens CEMT). SANTO DOMINGO DE LOS TSÁCHILAS: Otongachi, 960 m (3 specimens MQCAZ); Santo Domingo, 500 m (2 specimens MECN).

Literature records.

ESMERALDAS: 11 km SE San Lorenzo ( Edmonds 1994: 103); San Mateo ( Martínez and Pereira 1967: 68). LOS RÍOS: Quevedo, 45 m ( Edmonds 1994: 103). MANABÍ: 78 km NE de Chone, 450 m ( Edmonds 1994: 103). PICHINCHA [= SANTO DOMINGO DE LOS TSÁCHILAS]: 4 km SE de Santo Domingo, 500 m ( Edmonds 1994: 103). UNDETERMINED PROVINCE: without specific locality ( Balthasar 1939e: 242).

Temporal data.

Collected every month of the year.

Remarks.

Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 45-820 m a.s.l. Species was collected with pitfall traps baited with fungus, carrion, and human feces.