Molycria stanisici, Platnick & Baehr, 2006

Platnick, N. I. & Baehr, B., 2006, A Revision Of The Australasian Ground Spiders Of The Family Prodidomidae (Araneae: Gnaphosoidea), Bulletin of the American Museum of Natural History 2006 (298), pp. 1-287 : 47-63

Molycria stanisici , new species Figures 1, 103–107; Map 10

TYPES: Male holotype and female allotype taken in eucalypt forest at an elevation

of 80 m at Belmont Hills Bushlands , 27 ° 319S, 153 ° 079E, Queensland (male, pitfall trap, June 30–July 28, 2003, R. Raven; female, hand collected during day, Feb. 19, 2004, R. Raven), deposited in QMB (male S62261 ; female, S65673 ) .

ETYMOLOGY: The specific name is a patronym in honor of Dr. John Stanisic, Biodiversity Scientist at the Queensland Museum, and project leader of the Brisbane City Council invertebrate survey, during which the types were collected.

DIAGNOSIS: Males resemble those of M. mcleani in having a very small, sharp, narrow retrolateral tibial apophysis (fig. 105), but have a wider conductor tip (fig. 104); females have longitudinal, approximate median epigynal ducts (fig. 107).

MALE: Total length 3.36. Carapace 1.60 long, 1.40 wide, 0.62 high, length/width 1.14; sternum 0.90 long, 0.86 wide, length/width 1.05; abdomen 1.76 long, 1.02 wide; coxa I 0.60 long; relative length of coxae I–IV 1.00:0.86:0.73:0.93. Carapace pale yellow, with dark filigree net pattern; sternum yellow, with darker lateral margins; chelicerae grayish; endites, labium, legs pale; abdominal dorsum gray, with half moon-shaped pale spot in front of spinnerets, venter pale. Carapace weakly covered with gray, plumose setae. Eye group width 0.76 of caput width; AME 0.12; ALE 0.12; PME 0.14; PLE 0.12; AME–AME 0.06; AME–ALE 0.02; PME– PME 0.04; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.32; AME–AME 0.30; PME–PME 0.32. Clypeus 0.10 high. Abdomen covered with gray, plumose setae; ALS 0.47 of abdominal length, slightly separated, with heavily sclerotized transverse strip bearing strong macroseta at midline at origin. Tarsi I–IV ventrally with two rows of short, stout setae. Palp (figs. 103–105): conductor originating distally, broad, with greatly expanded, hood-shaped distal part, retrolaterally directed beak-shaped tip; median apophysis scythe-shaped, ventrally excavated; sperm duct weakly u-shaped; embolus long, thin, semicircular, embolar base broad, separated from tegulum; tibia about 1.5 times as long as wide, retrolateral tibial apophysis short, triangular.

FEMALE: Total length 2.94. Carapace 1.30 long, 1.22 wide, 0.52 high, length/width 1.07; sternum 0.84 long, 0.80 wide; abdomen 1.64 long, 0.96 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.76:0.72:0.96. Coloration as in male. Eye group width 0.77 of caput width; AME 0.13; ALE 0.10; PME 0.12; PLE 0.10; AME–AME 0.04; ALE–PLE 0.04; PME–PME 0.3. Clypeus 0.08 high. ALS 0.49 of abdominal length. Epigynum (figs. 106, 107): anterior margin with widely v-shaped projection, two medium sized, lateral, inverted u-shaped copulatory openings; epigynal ducts parallel along midline, forming basal triangles with spermathecae, spermathecae contiguous, sausage-shaped, twisted.

OTHER MATERIAL EXAMINED: Queensland: Beerwah Forestry Reserve , 26 ° 519S, 152 ° 579E, Oct. 10, 1990, heathland pitfall (M. Glover, QMB S32361 ), 1♀, Apr. 24, 1991, heathland pitfall (M. Glover, QMB S32362 ), 1♀ ; Belmont Hills Bushlands, site 1, elev. 80 m, 27 ° 319S, 153 ° 079E, Sept.1–Oct. 1, 2003, pitfall, eucalypt forest ( QMB S62385 ), 1 Oi, Oct. 30–Dec. 1, 2003, same ( QMB S65671 ), 2♀, Jan. 30–Mar. 1, 2004, same ( QMB S65672 ), 2 Oi , July 28–Sept. 1, 2003, same ( QMB S62218), 1 Oi , Jan. 2–29, 2004, same ( QMB S65670 ), 2 Oi ; Benarkin School , 26 ° 539S, 152 ° 099E, Jan. 26–Apr. 20, 1995, vine scrub pitfall (G. Monteith, QMB S46442 ), 1♀ ; Enoggera Reservoir , site 9, 27 ° 279S, 152 ° 549E, Jan. 27–Mar. 15, 2000, open forest pitfall (G. Monteith, J. Holt, QMB S65719), 1 Oi ; Gatton , 27 ° 349S, 152 ° 209E, Dec. 2001 – Jan. 2002, pitfall, soybean crop (S. Pearce, QMB S66690 ), 1♀ ; Gold Creek Reservoir, site 1, 27 ° 289S, 152 ° 529E, Sept. 1–Oct. 1, 2003, pitfall, spotted gum, open forest, elev. 140 m ( QMB S62416 ), 1♀ ; Hamilton , 27 ° 269S, 153 ° 049E, Sept. 9, 1973 (R. Raven, QMB S65716), 1 Oi ; Laidley Creek , 27 ° 319S, 152 ° 259E, Jan. 21, 1980, pitfall (M. Grant, QMB S65720 ), 2 Oi, 1♀ ; Moreton Island , 27 ° 119S, 153 ° 249E, Nov. 29, 1978, sweeping beach herbage (V. Davies, QMB S65717), 1♀ ; 3 km S Mount Deongwar, site 1, 27 ° 149S, 152 ° 159E, Oct. 14–Dec. 30, 1998, pitfall, wet sclerophyll forest, elev. 460 m (G. Monteith, D. Cook, QMB S49891 , 50622 ), 2 Oi , Dec. 30, 1998 – Mar. 26, 1999, same (G. Monteith, QMB S50374 ), 1♀ ; 3 km S Mount Deongwar , site 2, 27 ° 149S, 152 ° 159E, Oct. 14–Dec. 30, 1998, rainforest pitfall, elev. 450 m (G. Monteith, D. Cook, QMB S49898 ), 2♀ ; 3 km S Mount Deongwar , site 3, 27 ° 149S, 152 ° 169E, Oct. 14– Dec. 30, 1998, rainforest pitfall, elev. 450 m (G. Monteith, D. Cook, QMB S49895 ), 1♀ ; Ravensbourne National Park , 27 ° 229S, 152 ° 109E, Apr. 19, 2001, rainforest litter (M. Rix, QMB S67695 ), 1 Oi, 1♀ ; Woody Point, Brisbane , 27 ° 289S, 153 ° 039E, Jan. 20, 1988, in washing (H. Austin, QMB S3624 ), 1 Oi ; 3 km E Yarraman , 26 ° 509S, 151 ° 599E, pitfall (G., S. Monteith, QMB S65718 ), 1♀ .

DISTRIBUTION: Known only from southeastern Queensland (map 10).

Molycria monteithi , new species

Figures 108–112; Map 7

TYPES: Male holotype and female allotype taken in pitfall traps in open forest on the summit of Boat Mountain , 26 ° 099S, 151 ° 599E, Queensland (Dec. 14, 1994 – Jan. 26, 1995; G. Monteith), deposited in QMB (male S47790 , female S27930 ) .

ETYMOLOGY: The specific name is a patronym in honor of Dr. Geoff Monteith of the Queensland Museum, collector extraordinaire of many fascinating prodidomids.

DIAGNOSIS: This seems to be the northern sister species of M. stanisici , differing only in minor genitalic features: males have a smaller, retrolaterally uninflated conductor tip (fig. 109), and females have a projecting anterior epigynal margin (fig. 111) and longer median epigynal ducts (fig. 112).

MALE: Total length 2.88. Carapace 1.34 long, 1.22 wide, 0.46 high, length/width 1.10; sternum 0.86 long, 0.72 wide, length/width 1.19; abdomen 1.54 long, 0.80 wide; coxa I 0.52 long; relative length of coxae I–IV 1.00:0.92:0.69:0.96. Carapace orange, with dark filigree net pattern; sternum, chelicerae orange, grayish; endites, labium orange, distally pale; abdominal dorsum gray, with spear tip-shaped pale spot in front of spinnerets, venter pale; legs mottled with gray. Carapace weakly covered with gray, plumose setae. Eye group width 0.76 of caput width; AME 0.12; ALE 0.10; PME 0.12; PLE 0.10; AME–AME 0.04; AME–ALE 0.02; PME– PME 0.04; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.28; AME–AME 0.28; PME–PME 0.28. Clypeus 0.10 high. Abdomen covered with gray, plumose setae; ALS 0.56 of abdominal length, contiguous. Palp (figs. 108–110): conductor originating distally, broad, with greatly expanded, scytheshaped distal part, base of conductor, with large retrolateral, membranous lobe; median apophysis short, scythe-shaped, ventrally excavated; sperm duct weakly u-shaped; embolus long, thin, semicircular, embolar base broad, separated from tegulum; tibia about 1.5 times as long as wide, retrolateral tibial apophysis short, triangular.

FEMALE: Total length 2.90. Carapace 1.26 long, 1.22 wide, 0.50 high, length/width 1.03; sternum 0.80 long, 0.74 wide, length/ width 1.08; abdomen 1.64 long, 0.90 wide; coxa I 0.50 long; relative length of coxae I–IV 1.00:0.80:0.68:0.96. Coloration as in male. Eye eye group width 0.72 of caput width. Clypeus 0.08 high. ALS 0.53 of abdominal length. Epigynum (figs. 111, 112): anterior margin with widely v-shaped projection; epigynal ducts long, parallel along midline, forming basal triangles with spermathecae, spermathecae contiguous, sausage-shaped, highly coiled, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the type locality in southeastern Queensland (map 7).

Molycria mcleani , new species

Figures 113–117; Map 9

TYPES: Male holotype and female allotype taken in pitfall traps at an elevation of 40 m in scibbly gum and heath at Illawena State Forest, Drewvale , 27 ° 389S, 153 ° 049E, Queensland (July 29–Sept. 2, 2004; S. Wright), deposited in QMB (male S65695 , female S65696 ) .

ETYMOLOGY: The specific name is a patronym in honor of Mr. Stacey McLean, Senior Program Officer, Biodiversity Planning, Environment and Parks Branch, Brisbane City Council, who initiated the Brisbane habitat survey through which most of the specimens were found.

DIAGNOSIS: Males resemble those of M. stanisici but have a narrower conductor (fig. 114); females also resemble those of M. stanisici but have the median epigynal ducts anterolaterally divergent rather than approximate (fig. 117).

MALE: Total length 3.20. Carapace 1.54 long, 1.40 wide, 0.54 high, length/width 1.10; sternum 0.88 long, 0.82 wide, length/width 1.07; abdomen 1.66 long, 0.96 wide; coxa I 0.64 long; relative length of coxae I–IV 1.00:0.78:0.62:0.84. Carapace orange, with dark filigree net pattern; sternum orange, with darker lateral margins; chelicerae orange, grayish; endites, labium orange brown, distally pale; abdominal dorsum gray, with weak orange scutum, half moon-shaped pale spot in front of spinnerets, venter pale, epigastric area orange grayish; legs grayish orange. Carapace weakly covered with gray, plumose setae. Eye group width 0.7 of caput width; AME 0.15; ALE 0.14; PME 0.15; PLE 0.14; AME–AME 0.04; AME–ALE 0.02; PME–PME 0.02; PME–PLE 0.04; ALE– PLE 0.02; eye group AME–PME 0.34; AME–AME 0.34; PME–PME 0.32. Clypeus 0.10 high. Abdomen covered with gray, plumose setae; ALS 0.59 of abdominal length, slightly separated, with heavily sclerotized transverse strip bearing strong macroseta at midline origin. Palp (figs. 113–115): conductor originating distally, long, with greatly expanded, hood-shaped distal part, retrolaterally directed beak-shaped tip; median apophysis about twice as long as wide, ventrally excavated, with two distal tips; sperm duct weakly u-shaped; embolus long, thin, straight; tibia about 1.5 times as long as wide, retrolateral tibial apophysis short, triangular.

FEMALE: Total length 3.64. Carapace 1.30 long, 1.22 wide, 0.46 high, length/width 1.07; sternum 0.88 long, 0.82 wide, length/ width 1.07; abdomen 2.34 long, 1.10 wide; coxa I 0.60 long; relative length of coxae I–IV 1.00:0.83:0.76:0.93. Coloration as in male but without dorsal scutum. AME 0.14; ALE 0.12; PME 0.14; PLE 0.12; AME–AME 0.02; PME–PME 0.04; eye group AME– PME 0.30; AME–AME 0.30; PME–PME 0.30. Clypeus 0.08 high. ALS 0.51 of abdominal length. Epigynum (figs. 116, 117): anterior margin with sharply pointed projection; epigynal ducts in x-shaped position with spermathecae, spermathecae contiguous, sausage-shaped, twisted.

OTHER MATERIAL EXAMINED: Queensland: Buhot Creek, Burbank, 27 ° 369S, 153 ° 109E, Dec. 1, 2003 – Jan. 1, 2004, riparian forest, pitfall, elev. 50 m ( QMB S67110) 2 Oi , Nov. 6, 2003, hand collected, same ( QMB S67108 ), 1 Oi ; Camerons Scrub, top ridge, knoll top, 27 ° 309S, 152 ° 449E, Sept. 15, 1998 – Jan. 13, 1999, vine scrub pitfall, elev. 90–160 m (G. Monteith, D. Cook, G. Thompson, QMB S49892 , 50602 , 50617 , 51143 ), 2 Oi , 2♀, Jan. 13–May 16, 1999, vine scrub pitfall, elev. 90–110 m (G. Monteith, QMB S50607, 52448 ), 4♀ ; Camerons Scrub, snig track, 27 ° 319S, 152 ° 449E, Sept. 15–Nov. 11, 1998, vine scrub pitfall, elev. 120 m (G. Monteith, D. Cook, G. Thompson, QMB S51135 ), 2 Oi , 1♀, Nov. 11, 1998 – Jan. 13, 1999, same ( QMB S50609 ), 1 Oi ; Dutton Park , 27 ° 299S, 153 ° 019E, July 1, 1995, in silk sac under pan on ground under house (P. Lawless, QMB S50584 ), 1♀ ; 3 mi E Hatton Vale, 27 ° 349S, 152 ° 289E, Nov. 24, 1962, elev. 150 m (E. Ross, D. Cavagnaro, CAS), 1♀ ; Illawena State Forest, Drewvale, 27 ° 389S, 153 ° 049E, July 1–29, 2003, scibbly gum, heath, pitfall, elev. 40 m (S. Wright, QMB S62668 ), 2 Oi, Oct. 2–31, 2003, same ( QMB S65666 ), 1♀, Jan. 2–30, 2004, same ( QMB S65661 ), 1 Oi, 1 Oi, Jan. 30–Mar. 1, 2004, same ( QMB S65668 ), 3 Oi, 1♀, Feb. 17, 2004, same but hand collected ( QMB S67109 , 67111 ), 1 Oi, 3♀, Mar. 1–31, 2004, same ( QMB S65663 ), 1♀, July 29–Sept. 2, 2004, same (S. Wright, QMB S62179), 1 Oi, 3♀, Karawatha Forest , 27 ° 379S, 153 ° 059E, Dec, 17, 1994– Apr. 23, 1995, melaleuca forest pitfall (D. Stewart, QMB S65700 ), 1♀ ; Karawatha Forest, site 6, 27 ° 379S, 153 ° 059E, July 1–29, 2003, pitfall, eucalypt woodland, elev. 60 m ( QMB S62912 ), 1 Oi ; The Knobby, 27 ° 309S, 152 ° 359E, Sept. 15– Nov. 11, 1998, vine scrub pitfall, elev. 240 m (G. Monteith, D. Cook, G. Thompson, QMB S51133 ), 4 Oi , Nov. 11, 1998 – Jan. 13, 1999, same ( QMB S50620 ), 1♀ ; Mount Cotton , 27 ° 379S, 153 ° 139E, Sept. 3–Dec. 12, 1997, rainforest pitfall, elev. 200 m (G. Monteith, QMB S32471 ), 1♀ ; North Stradbroke Island, Enterprise, Scribbly Gum # 3, 27 ° 369S, 153 ° 279E, Jan. 10, 2002, hand collecting, day, elev. 70 m ( QMB S56495 ), 1♀ ; North Stradbroke Island, Enterprise, Scribbly Gum # 2, 27 ° 379S, 153 ° 279E, Jan. 8–22, 2002, pitfall, elev. 120 m ( QMB S55567 ), 1 Oi , Jan. 10, 2002, hand collecting, night, elev. 120 m ( QMB S56494), 1♀ ; North Stradbroke Island, Enterprise, Blackbutt # 3, 27 ° 359S, 153 ° 289E, Jan. 8–22, 2002, pitfall, elev. 80 m ( QMB S55578), 3 Oi , 1♀ ; North Stradbroke Island, Ibis Alpha, 27 ° 389S, 153 ° 269E, open forest, June 1997 (U. Nolte, QMB S40957 ), 1 Oi ; North Stradbroke Island, Yarraman , 27 ° 279S, 153 ° 309E, Mar.-Sept. 1997, open forest (U. Nolte, QMB S41003 , 41007 , 41011 ), 1 Oi, 3♀ ; Perry’s Knob, 27 ° 369S, 152 ° 369E, Sept. 15–Nov. 11, 1998, vine scrub pitfall, elev. 200 m (G. Monteith, D. Cook, G. Thompson, QMB S51139 ), 2 Oi , 2♀, Nov. 11, 1998 – Jan. 13, 1999, same ( QMB S49901 , 50608 ), 4 Oi, 1♀, Jan. 13–May 16, 1999, same (G. Monteith, QMB S52461 ), 2 Oi, 3♀ ; Rochedale State Forest, Brisbane , 27 ° 379S, 153 ° 099E, Feb. 14–27, 1980, pitfall (V. Davies, R. Raven, QMB S65699 ), 1♀, Apr. 2, 1980, pitfall (V. Davies, R. Raven, QMB S65697 ), 1♀ ; Teviot Brook , 27 ° 519S, 152 ° 579E, Oct. 21, 1973, under stone (V. Davies, QMB S65698 ), 1♀ ; Toowoomba , 27 ° 349S, 151 ° 579E, Aug. 19, 1997, in house (B. Scholz, QMB S35406 ), 1♀ .

DISTRIBUTION: Known only from southeastern Queensland (map 9).

Molycria burwelli , new species

Figures 118–122; Map 11

TYPES: Male holotype and female allotype taken in rainforest at an elevation of 110 m at Enoggera Reservoir , 27 ° 279S, 152 ° 559E, Queensland (male, Aug. 7–Oct. 16, 1999, pitfall, G. Monteith, J. Holt; female, Mar. 15–May 18, 2000, malaise trap, C. Burwell, S. Evans), deposited in QMB (male S51638, female S39612 ) .

ETYMOLOGY: The specific name is a patronym in honor of Dr. Chris Burwell of the Queensland Museum, one of the collectors of the allotype.

DIAGNOSIS: Males of this distinctive species can easily be recognized by the long, semicircular embolus and wide conductor tip and the two membranous, spatulate projections at the retrolateral base of the conductor (fig. 119), females by the arched, projecting anterior epigynal margin (fig. 121).

MALE: Total length 3.74. Carapace 1.66 long, 1.56 wide, 0.40 high, length/width 1.06; sternum 1.02 long, 0.92 wide, length/width 1.11; abdomen 2.08 long, 1.00 wide; coxa I 0.66 long; relative length of coxae I–IV 1.00:0.88:0.73:0.94. Carapace orange, with dark filigree net pattern; sternum orange, with darker lateral margins; chelicerae grayish; endites, labium orange brown, distally pale; abdominal dorsum gray, with spear tipshaped pale spot in front of spinnerets, venter pale; legs mottled with gray. Carapace weakly covered with cinnamon, plumose setae. AME elevated, eye group width 0.77 of caput width; AME 0.15; ALE 0.14; PME 0.14; PLE 0.14; AME–AME 0.04; AME– ALE 0.02; PME–PME 0.06; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.34; AME–AME 0.34; PME–PME 0.34. Clypeus 0.08 high. Abdomen covered with cinnamon, plumose setae; ALS 0.57 of abdominal length, contiguous. Palp (figs. 118–120): conductor originating distally, broad, with greatly expanded distal part, retrolaterally directed tip; base of conductor with two retrolateral, membranous, spatulate projections; median apophysis tiny, hooked, ventrally excavated, with two distal tips; embolus long, thin, semicircular, embolar base broad, separated from tegulum; tibia about 1.5 times as long as wide, retrolateral tibial apophysis short, triangular.

FEMALE: Total length 4.84. Carapace 1.64 long, 1.62 wide, 0.50 high, length/width 1.01; sternum 1.06 long, 0.96 wide, length/ width 1.10; abdomen 3.20 long, 2.00 wide; coxa I 0.64 long; relative length of coxae I–IV 1.00:0.75:0.72:0.97. Coloration as in male. Eye eye group width 0.74 of caput width; AME 0.16; PME 0.15; PME–PME 0.04; eye group AME–PME 0.36; AME–AME 0.36. ALS 0.45 of abdominal length. Epigynum (figs. 121, 122): atrium with wide arched anterior margin, sharply pointed projection; two copulatory openings, forming cup; epigynal ducts short, spermathecae contiguous, sausage-shaped, twisted, in horizontal position.

OTHER MATERIAL EXAMINED: Queensland: Cooloola, 26 ° 129S, 153 ° 039E, Apr. 1978, pitfall (V. Davies, R. Raven, QMB S65728), 1 Oi , 4♀, Oct. 1978, high dune pitfall (K. Plowman, QMB S65729 ), 1 Oi ; Cooloola National Park , 25 ° 579S, 153 ° 079E, Jan. 8–Mar. 3, 1992, rainforest pitfall, elev. 140 m (D. Cook, QMB S25212 ), 1♀ ; Gallangowan , 26 ° 269S, 152 ° 179E, Nov. 10– Dec. 29, 1974, rainforest pitfall, elev. 487 m (G., S. Monteith, QMB S65730 ), 1 Oi ; Lake Poona, Cooloola National Park , 26 ° 129S, 153 ° 039E, Apr. 1978 (G. Monteith, QMB S65723 ), 1♀ ; Little Yabba Creek , 26 ° 379S, 152 ° 419E, Jan. 7– Mar. 2, 1992, rainforest pitfall, intercept, elev. 150 m (D. Cook, QMB S30317 , 30318 ), 2 Oi ;

Mount Glorious , 27 ° 209S, 152 ° 469E, Dec. 21, 1.00:0.93:0.82:0.96. Carapace orange, mottled 1970, rainforest litter (H. Williams, QMB with gray; sternum orange, with darker lateral S15642 ), 1♀, Dec. 1981 (A. Hiller, QMB margins; chelicerae grayish; endites, labium S65725), 1♀, Dec. 14, 1981 – Mar. 5, 1982, orange, distally pale ; abdominal dorsum gray, rainforest malaise trap (A. Hiller, QMB

with spear tip-shaped pale spot in front of S65727 ), 1♀, Dec. 21, 1982 – Jan. 13, 1983,

malaise trap (A. Hiller , QMB S65726 ), 1♀, spinnerets, venter grayish ; legs orange-brown, Hiller property, Dec. 12, 1998 – Jan. 28, 1999, femora grayish. Carapace weakly covered with malaise (N. Power, QMB S52460 ), 1 Oi ; Mount cinnamon, plumose setae. Eye group width Nebo, 27 ° 239S, 152 ° 479E, Oct. 16, 1978, mixed 0.83 of caput width ; AME 0.12; ALE 0.12; sclerophyll pitfall (A. Rozefelds , QMB S65731 ), PME 0.14 ; PLE 0.12; AME–AME 0.04; 1♀, Nov. 6, 1978, under plastic sheeting near AME–ALE 0.02; PME–PME 0.02; PME– house, and pitfall (A. Rozefelds , QMB S65724), PLE 0.04 ; ALE–PLE 0.02; eye group AME– 2♀; Searys Scrub , Cooloola National Park, PME 0.28 ; AME–AME 0.28; PME–PME

26

°

129S, 153

°

039E, Feb. 6, 1976, in or under

0.30. Clypeus 0.08 high. Abdomen covered bark (R. Raven, V. Davies, QMB S65733 ), 1Oi,

with cinnamon, plumose setae; ALS 0.6 of 2♀, Feb. 3–7, 1976, pitfall (R. Raven, V. Davies,

QMB S65732 ), 1 Oi ; Stony Creek, via Samford, abdominal length, slightly separated. Palp 27 ° 209S, 152 ° 489E, Feb. 2–Apr. 8, 1995, rain- (figs. 123–125): conductor originating distally, forest pitfall (G. Monteith, H. Janetzki, QMB broad, ventrally folded distal end greatly S34502 ), 1 Oi ; Sunday Creek , 26 ° 409S, 152 ° 349E, expanded, tip arrow-shaped, retrolaterally di- Dec. 18, 1996 – Jan. 20, 1997, rainforest pitfall rected ; median apophysis tiny, hooked, ven- (G. Monteith, QMB S35374 ), 1 Oi. trally excavated ; sperm duct weakly u-shaped; DISTRIBUTION: Known only from south- embolus long, thin, semicircular ; embolar base eastern Queensland (map 11). broad, separated from tegulum; tibia about 1.5

times as long as wide, retrolateral tibial

Molycria thompsoni , new species apophysis short, rectangular.

Figures 123–127; Map 7 FEMALE: Total length 4.48. Carapace TYPE: Female holotype taken in a pitfall 1.60 long, 1.58 wide, 0.56 high, length/width trap in rainforest at Lower Dry Creek,

Kroombit Tops, site 9, 45 km SSW Calliope,

24 ° 249S, 151 ° 019E, Queensland (Dec. 11–18,

1983; G. Monteith, V. Davies, J. Gallon, G.

Thompson), deposited in QMB ( S65736 ).

ETYMOLOGY: The specific name is a pa-

tronym in honor of Geoff Thompson of the

Queensland Museum, one of the collectors of

the holotype.

DIAGNOSIS: Males resemble those of M.

burwelli but have an arrow-shaped conductor

tip and shorter median tegular apophysis

(fig. 124); females resemble those of M. amphi

in having a triangular epigynal atrium, but

have the atrium much longer, extending fully

to the spermathecae (figs. 126, 127).

MALE: Total length 3.10. Carapace 1.40

long, 1.34 wide, 0.46 high, length/width 1.04;

sternum 0.90 long, 0.82 wide, length/width Map 11. Circle, Molycria burwelli , new species. 1.10; abdomen 1.70 long, 0.90 wide; coxa I Square, Molycria cooki , new species. Triangle, 0.56 long; relative length of coxae I–IV Molycria raveni , new species.

+ Figs. 123–127. Molycria thompsoni , new species. 123. Left male palp, prolateral view. 124. Same, ventral

view. 125. Same, retrolateral view. 126. Epigynum, ventral view. 127. Same, dorsal view.

1.01; sternum 1.00 long, 0.90 wide, length/ width 1.11; abdomen 2.88 long, 1.68 wide; coxa I 0.64 long; relative length of coxae I–IV 1.00:0.84:0.72:0.97. Coloration as in male. Eye group width 0.82 of caput width; AME 0.14; PME–PLE 0.06; eye group AME–PME 0.32; AME–AME 0.32; PME–PME 0.34. ALS 0.43 of abdominal length. Epigynum (figs. 126, 127): atrium roughly triangular; epigynal ducts long, in v-shaped position, spermathecae contiguous, sausage-shaped, twisted.

OTHER MATERIAL EXAMINED: Queensland: 0.5 km S Fairlies Knob, 25 ° 319S, 152 ° 179E, July 21–Oct. 20, 2000, vine scrub pitfall, elev. 300 m (D. Cook, S. Wright, E. Vanderduys, QMB S61037 , S65734 , S66635 ), 4 Oi , 4♀, Dec. 20, 2000 – Mar. 23, 2001, hoop pine scrub pitfall (D. Cook, G. Monteith, QMB S65735), 1♀ ; 3.5 km SE Fairlies Knob , 25 ° 329S, 152 ° 199E, July 21– Oct. 20, 2000, vine scrub pitfall, elev. 120 m (D. Cook, S. Wright, E. Vanderduys, QMB), 1♀ ; Mount Goonaneman, near Childers , 25 ° 269S, 152 ° 089E, Nov. 3–7, 1980, elev. 670 m (V. Davies, R. Raven, QMB S65738), 1 Oi ; 1 km N Mount Walsh , 25 ° 349S, 152 ° 039E, June 26–Oct. 9, 1999, vine scrub pitfall, elev. 320 m (D. Cook, QMB S65737), 1 Oi .

DISTRIBUTION: Known only from mideastern Queensland (map 7).

Molycria bulburin , new species

Figures 128–132; Map 8

TYPES: Male holotype and female allotype taken in pitfall traps in rainforest at an elevation of 609 m on the Bulburin Plateau, via Miriamvale , Bulburin State Forest , 24 ° 309S, 151 ° 359E, Queensland (June 1–Oct. 5, 1974; G., S. Monteith), deposited in QMB (male S65722 , female S65721 ) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males have a greatly expand- ed, scythe-shaped conductor tip (fig. 129), females have a prominent median protuberance on the anterior epigynal margin that reaches almost to a median ledge (fig. 131).

MALE: Total length 3.40. Carapace 1.52 long, 1.50 wide, 0.60 high, length/width 1.01; sternum 0.94 long, 0.80 wide, length/width 1.17; abdomen 1.88 long, 1.20 wide; coxa I 0.60 long; relative length of coxae I–IV 1.00:0.80:0.73:0.90. Carapace orange, with dark filigree net pattern; sternum orange; chelicerae grayish; endites, labium orange, distally pale; abdominal dorsum gray, with spear tip-shaped pale spot in front of spinnerets, venter grayish, epigastric area orange; legs orange, mottled with gray. Carapace weakly covered with cinnamon, plumose setae. Eye group width 0.77 of caput width; AME 0.14; ALE 0.14; PME 0.14; PLE 0.12; AME–AME 0.04; AME–ALE 0.02; PME–PME 0.04; PME–PLE 0.04; ALE– PLE 0.02; eye group AME–PME 0.34; AME–AME 0.32; PME–PME 0.32. Clypeus 0.10 high. Abdomen covered with cinnamon, plumose setae; ALS 0.59 of abdominal length, contiguous. Palp (figs. 128–130): cymbium with retrobasal hook; conductor originating distally, broad, with greatly expanded, scythe-shaped distal part, retrolaterally directed tip; median apophysis wide, ventrally excavated; sperm duct straight; embolus long, thin, semicircular, embolar base broad, separated from tegulum; tibia about 1.8–2.0 times as long as wide, retrolateral tibial apophysis small, conical.

FEMALE: Total length 3.88. Carapace 1.64 long, 1.60 wide, 0.44 high, length/width 1.02; sternum 1.00 long, 0.86 wide, length/ width 1.16; abdomen 2.24 long, 1.22 wide; coxa I 0.64 long; relative length of coxae I–IV 1.00:0.84:0.75:0.97. Coloration as in male. Eye group width 0.74 of caput width; AME 0.15; ALE 0.12; AME–AME 0.02. ALS 0.61 of abdominal length. Epigynum (figs. 131, 132): atrium with v-shaped, prominent medi- an projection on anterior margin reaching almost to v-shaped median ledge; epigynal ducts short, parallel, spermathecae less than their diameter apart, sausage-shaped, twist- ed, in horizontal position.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Known only from the Bulburin Plateau, southeastern Queensland (map 8).

Molycria nipping , new species Figures 133–137; Map 8

TYPES: Male holotype and female allotype taken by pyrethrum fogging of trees and logs in a vine scrub forest at an elevation of 200 m at Nipping Gulley , site 2, 25 ° 409S, 151 ° 269E, Queensland (Oct. 9, 1998; G. Monteith), deposited in QMB (male S47757 , female S67183 ) .

ETYMOLOGY: The specific name is a noun females have long lateral epigynal margins in apposition taken from the type locality. extending medially (fig. 136).

DIAGNOSIS: Males have a distinctively MALE: Total length 3.26. Carapace 1.44 small conductor tip (fig. 134) and a short, long, 1.34 wide, 0.42 high, length/width 1.07; wide retrolateral tibial apophysis (fig. 135); sternum 0.90 long, 0.78 wide, length/width 1.15; abdomen 1.82 long, 0.42 wide; coxa I 0.54 long; relative length of coxae I–IV 1.00:0.81:0.74:0.96. Carapace orange, mottled with gray, sternum orange, chelicerae, legs grayish; endites, labium grayish, distally pale; abdominal dorsum gray, with weak orange scutum, half ring-shaped pale spot in front of spinnerets, venter pale gray. Carapace weakly covered with gray, plumose setae. AME elevated, eye group width 0.71 of caput width; AME 0.15; ALE 0.10; PME 0.13; PLE 0.10; AME–AME 0.02; AME–ALE 0.02; PME– PME 0.04; PME–PLE 0.06; ALE–PLE 0.02; eye group AME–PME 0.34; AME–AME 0.32; PME–PME 0.30. Clypeus 0.10 high. Abdomen covered with gray, plumose setae; ALS 0.55 of abdominal length, contiguous. Palp (figs. 133–135): conductor originating distally, straight, with shallow groove; median apophysis scythe-shaped, with long median tip; embolus finger-shaped, with s-shaped sharp tip; tibia about 1.8–2.0 times as long as wide, retrolateral tibial apophysis rectangular, with distally bent tip.

FEMALE: Total length 3.66. Carapace 1.36 long, 1.28 wide, 0.38 high, length/width 1.06; sternum 0.88 long, length/width 1.13; abdomen 2.30 long, 1.36 wide; coxa I 0.52 long; relative length of coxae I–IV 1.00:0.88:0.81:1.00. Coloration as in male but without dorsal scutum. Eye group width 0.81 of caput width; AME 0.13; PME 0.12; eye group AME–PME 0.28; AME–AME 0.28; PME–PME 0.28. Clypeus 0.08 high. ALS 0.44 of abdominal length. Epigynum (figs. 136, 137): atrium with long lateral margins extending medially; epigynal ducts parallel along midline, anterior part recurved, spermathecae nearly contiguous, oval, twist- ed, in horizontal position, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: Queensland: Gurgeena Plateau, 25 ° 279S, 151 ° 239E, Aug. 22–Oct. 10, 1998, rainforest pitfall (G. Monteith, QMB S47753 ), 1 Oi, Jan. 27–June 2, 1999, rainforest pitfall (G. Monteith, G. Thompson, QMB S68058), 1 Oi , 1♀ ; Nipping Gulley, site 1, 25 ° 419S, 151 ° 269E, Aug. 21–Oct. 9, 1998, pitfall, elev. 200 m (G. Monteith, QMB S47752 ), 5 Oi , Oct. 9–Dec. 18, 1998, rainforest pitfall, elev. 200 m (G. Monteith, C. Gough, QMB S47761 ), 1♀, Dec. 18, 1998 – Jan. 26, 1999, rainforest pitfall, elev. 300 m (G. Monteith, C. Gough, QMB S49896 ), 2 Oi, Jan. 26–June 2, 1999, same (G. Monteith, G. Thompson, QMB S52441 ), 1 Oi ; Nipping Gulley , site 2, 25 ° 419S, 151 ° 269E, Jan. 25–June 2, 1999, rainforest pitfall, elev. 300 m (G. Monteith, G. Thompson, QMB S52446), 1♀ ; Nipping Gulley, site 3, 25 ° 419S, 151 ° 259E, Aug. 21–Oct. 9, 1998, open forest pitfall, elev. 150 m (G. Monteith, QMB S47807 ), 3 Oi , 1♀ ; Nipping Gulley, site 5, 25 ° 429S, 151 ° 269E, Dec. 18, 1998 – Jan. 26, 1999, rainforest pitfall, elev. 240 m (G. Monteith, C. Gough, QMB S49893), 1 Oi, 1♀, Jan. 25–June 2, 1999, same (G. Monteith, G. Thompson, QMB S52440 ), 1 Oi , 3♀ .

DISTRIBUTION: Known only from southeastern Queensland (map 8).

Molycria dawson , new species

Figures 138–142; Map 9

TYPES: Male holotype and female allotype taken in pitfall traps (7, 8) in an open forest along Dawson River at Nathan Gorge , 25 ° 279S, 150 ° 089E, Queensland (Sept. 12– Nov. 13, 1996; P. Lawless), deposited in QMB (male S66624 , female S66625 ) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Males resemble those of M. nipping in having a ventrally expanded tip of the palpal tibia, but have a low, hirsute ventral enlargement at the base of the palpal femur and a short, wide retrolateral tibial apophysis well separated from a dorsal patch of stiff setae (fig. 140); females have a distinctive epigynum, with a small atrium bearing a narrow v-shaped anterior projection (fig. 141) and approximate epigynal ducts (fig. 142).

MALE: Total length 3.18. Carapace 1.38 long, 1.34 wide, 0.48 high, length/width 1.03; sternum 0.90 long, 0.78 wide, length/width 1.15; abdomen 1.80 long, 0.94 wide; coxa I 0.58 long; relative length of coxae I–IV 1.00:0.82:0.69:0.89. Carapace, sternum, chelicerae grayish orange, endites, labium orange +

Figs. 133–137. Molycria nipping , new species. 133. Left male palp, prolateral view. 134. Same, ventral

view. 135. Same, retrolateral view. 136. Epigynum, ventral view. 137. Same, dorsal view.

brown, distally pale; abdominal dorsum gray, with weak orange scutum, spear tip-shaped pale spot in front of spinnerets, venter pale; legs orange. Carapace weakly covered with shiny, plumose setae. Eye group width 0.75 of caput width; AME 0.14; ALE 0.10; PME 0.14; PLE 0.12; AME–AME 0.02; AME– ALE 0.02; PME–PME 0.04; PME–PLE 0.04; ALE–PLE 0.02; eye group AME–PME 0.32; AME–AME 0.30; PME–PME 0.32. Clypeus 0.10 high. Abdomen covered with gray, plumose setae; ALS 0.51 of abdominal length, slightly separated. Palp (figs. 138– 140): apophyses and embolus situated on distal half of tegulum; conductor originating distally, broad, groove shallow; median apophysis scythe-shaped, ventrally excavat- ed, with long median tip; sperm duct ushaped; embolus short, thin, sinuous, embolar base separated from tegulum, lobate; tibia about 1.5 times as long as wide, dorsally with field of short, stiff setae, retrolateral tibial apophysis short, rectangular; femur with ventral process.

FEMALE: Total length 4.04. Carapace 1.44 long, 1.38 wide, 0.50 high, length/width 1.09; sternum 0.88 long, 0.80 wide, length/ width 1.10; abdomen 2.60 long, 1.32 wide; coxa I 0.56 long; relative length of coxae I–IV 1.00:0.75:0.64:0.96. Coloration as in male but without dorsal scutum. Eye group width 0.83 of caput width; AME 0.13; PLE 0.10; AME– AME 0.04; eye group AME–PME 0.30. ALS 0.41 of abdominal length. Epigynum (figs. 141, 142): atrium small, anterior margin with narrow v-shaped projection; epigynal ducts short, parallel along midline, spermathecae contiguous, sausage-shaped, twist- ed, with one dorsal diverticulum.

OTHER MATERIAL EXAMINED: Queensland: Cabbage Tree Creek, Taroom , 25 ° 279S, 150 ° 019E, Sept. 13, 1996 (P. Lawless, QMB S36464 ), 1 Oi ; Dawson River at Nathan Gorge , traps 3, 4, 25 ° 279S, 150 ° 089E, Sept. 12–Nov. 13, 1996, riverine rainforest pitfall (P. Lawless, QMB S30937 ), 8 Oi ; Dawson River at Nathan Gorge , traps 6, 7, 25 ° 279S, 150 ° 089E, Sept. 12– Nov. 13, 1996, riverine rainforest pitfall (P. Lawless, QMB S30935 ), 1♀ ; Dawson River at Nathan Gorge , traps 7, 8, 25 ° 279S, 150 ° 089E, Sept. 12–Nov. 13, 1996, open forest pitfall (P. Lawless, QMB S30934 ), 3 Oi ; Hurdle Gully , 13 km WSW Monto, 24 ° 519S, 151 ° 009E, Dec. 20, 1997 – Apr. 26, 1998, vine scrub pitfall, elev. 350 m (G. Monteith, QMB S42456 , 42457 ), 2♀ ; Kroombit Tops, Barracks, site 7, 45 km SSW Calliope , 24 ° 259S, 151 ° 039E, open forest pitfall, Dec. 10–18, 1983 (G. Monteith, V. Davies, J. Gallon, G. Thompson, QMB S66628 ), 1 Oi, 1♀ ; Kroombit Tops, Northern Escarpment , 45 km SSW Calliope, site 11, 24 ° 229S, 151 ° 019E, Dec. 11–18, 1983, open forest pitfall (G. Monteith, V. Davies, J. Gallon, G. Thompson, QMB S66627 ), 1 Oi ; Kroombit Tops, Upper Dry Creek , 45 km SSW Calliope, 24 ° 259S, 151 ° 039E, Dec. 9–19, 1983, open forest (V. Davies, J. Gallon, QMB S66626 ), 1♀ ; 3 km NNE Mount Bassett , 26 ° 259S, 148 ° 559E, Dec. 13, 2001 – Mar. 5, 2002, vine scrub pitfall, elev. 520 m (G. Monteith, D. Cook, QMB S62437 ), 1 Oi ; 0.5 km NW Mount Beecher , 23 ° 559S, 151 ° 119E, Dec. 15, 1999 – Mar. 20, 2000, vine scrub pitfall, elev. 20 m (G. Monteith, QMB S57265 ), 4♀ ; 1 km S Mount Gavial, 23 ° 369S, 150 ° 299E, Dec. 17, 1998 – Mar. 14, 1999, open forest pitfall, intercept, elev. 400 m (D. Cook, QMB S50604 , 50605 ), 1 Oi , 2♀, Mar. 14–June 28, 1999, same ( QMB S58557), 1♀ ; 3 km SSE Mount Gavial , 23 ° 379S, 150 ° 299E, Dec. 18, 1998 – Mar. 14, 1999, open forest intercept, elev. 450 m (D. Cook, QMB S50603 ), 2 Oi ; Nathan Gorge, Taroom District, 25 ° 279S, 150 ° 089E, Sept. 12–Nov. 13, 1996, riverine rainforest pitfall (P. Lawless, QMB S30941 ), 1 Oi , Nov. 13, 1996 – Jan. 13, 1997, same ( QMB S37112 ), 3 Oi, 2♀ ; Orchid Beach, Fraser Island, 24 ° 589S, 153 ° 199E, Mar. 7–Oct. 1, 1996, pitfall, heathland with banksias, eucalypts (R. Raven, QMB S31273 ), 1 Oi , 2♀, Aug. 20–21, 1997, night collecting (P. Lawless, QMB S38872 , 38892 ), 1Oi, 2♀, Aug. 20–Dec. 17, 1997, pitfall (R. Raven, Fishburn, P. Lawless, QMB S41775 , 43448 , 43463 , 51207 ), 7Oi, 8♀ ; Taroom District, Boggomoss No. 12, Nathan Gorge , 25 ° 279S, 150 ° 089E, Nov. 13, 1996 – Jan. 13, 1997, riverine rainforest pitfall (P. Lawless, QMB S36708 ), 2 Oi, pitfall (D. Cook, G. Monteith, QMB S36347), 1♀ ; Taroom, Boggomoss Station , 25 ° 259S, 150 ° 019E, Nov. 11, 1996 – Jan. 13, 1997, pitfall, ferns, eucalypts (P. Lawless, H. Janetzki, D. Cook, QMB S36781 ), 1♀ .

DISTRIBUTION: Known only from eastern Queensland (map 9).