Mangelia costulata Risso 1826

Öztürk, Bilal, 2021, Mangelia (Gastropoda, Conoidea) species of the Turkish coasts with description of Mangelia vanaartseni sp. nov., Zootaxa 4933 (2), pp. 241-262 : 248

Mangelia costulata Risso 1826

( Fig. 6 )

[= Mangelia smithii ( Forbes 1840) ]

Materials: 07.07.1999, Yumurtalık (Iskenderun Bay), Levantine Sea , sandy mud, 26 m, 1 spm ( ESFM-GAS/1999- 15 ) ; 05. 10.2001, Nemrut Bay (Aliağa), Aegean Sea , mud, 15 m, 1 spm ( ESFM-GAS/2001-41 ) ; 21.10.2001, Salih Island (Bodrum), Aegean Sea , P. oceanica + sand, 13 m, 5 spm ( ESFM-GAS/2001-42 ) ; 01.05.2003, Ildır Bay, Aegean Sea , 45 m, sand, 1 spm ( ESFM-GAS/2003-102 ) ; 09.06.2003, Ildır Bay, Aegean Sea , sand, 50 m, 1 spm + 1 sh. ( ESFM-GAS/2003-103 ) ; 23.09.2005, Anamur, Levantine Sea , muddy sand, 50 m, 2 sh. ( ESFM-GAS/2005- 310 ) ; 27.09.2005, Antalya Bay, Levantine Sea , muddy sand, 25 m, 1 sh. ( ESFM-GAS/2005-308 ) ; 03.10.2005, Kaş, Levantine Sea , P. oceanica , 9 m, 2 spm ( ESFM-GAS/2005-101 ) ; 06.10.2005, Fethiye Bay, Levantine Sea , sandy mud with Flabellia petiolata (green alga) fragments, 25 m, 1 sh. ( ESFM-GAS/2005-309 ) ; 07.10.2005, Göçek, Le-vantine Sea , Zostera spp. (eelgrass), 3 m, 1 spm ( ESFM-GAS/2005-307 ) ; 15.07.2007, Iskenderun Bay, Levantine Sea , muddy sand + Halophila sp. (sea grass), 11 m, 2 spm ( ESFM-GAS/2007-15 ) ; 20.12.2008, Avşa Island, Sea of Marmara , sandy mud, 19 m, 2 spm ( ESFM-GAS/2008-17 ) ; 08.06.2013, Sea of Marmara , sand, 10 m, 1 spm + 3 sh. ( ESFM-GAS/2013-85 ) ; 09.06.2013, Sea of Marmara , sand, 10 m, 1 spm+ 1sh. ( ESFM-GAS/2013-84 ) ; 10.06.2013, Sea of Marmara , gravelly sandy mud with shell fragments, 25 m, 1 sh. ( ESFM-GAS/2013-88 ) ; 12.06.2013, Sea of Marmara , muddy sand with shell fragments, 25 m, 4 spm + 3 sh. ( ESFM-GAS/2013-87 ) ; 13.06. 2013, Sea of Marmara , sandy mud with shell fragments, 10 m, 5 spm + 2 sh. ( ESFM-GAS/2013-32 ) ; 14.06.2013, Sea of Marmara , sand with shell fragments, 10 m, 1 spm ( ESFM-GAS/2013-48 ) ; 15.06.2013, Sea of Marmara , sand, 10 m, 4 spm ; 17.06.2013, Sea of Marmara , sandy mud, 10 and 25 m, 3 spm +7 sh. ( ESFM-GAS/2013-68 ) ; 18.06.2013, Sea of Marmara , mud with shell fragments, 10 m, 2 spm ( ESFM-GAS/2013-86 ) ; 20.06.2013, Sea of Marmara , mud with shell fragments, 50 and 100 m, 5 spm ( ESFM-GAS/2013-81 ) ; 21.06.2013, Sea of Marmara , mud with shell fragments, 50 and 200 m, 1spm+1 sh. ( ESFM-GAS/2013-82 ) ; 24.06.2013, Sea of Marmara , mud, 50 m, 1 spm ( ESFM- GAS/2013-83 ) ; 26.06.2013, Sea of Marmara , sandy mud with shell fragments, 25 and 100 m, 4 spm + 1 sh ( ESFM- GAS/2013-33 ) ; 11.04.2014, G̹ll̹k Bay, Aegean Sea , muddy sand, 45 m, 2 sh. ( ESFM-GAS/2014-48 ) ; 24.07.2014, Akköy, Aegean Sea , mud, 67 m, 1 sh. ( ESFM-GAS/2014-50 ) ; 25.07.2014, Güllük Bay, Aegean Sea , sandy mud, 49 m, 2 sh. ( ESFM-GAS/2014-52 ) ; 26.07.2014, G̹ll̹k Bay, Aegean Sea , sandy mud with shell fragments, 46 m, 1 spm ( ESFM-GAS/2014-51 ) ; 25.10.2014, Ildır Bay, Aegean Sea , sandy mud, 64 m, 5 spm + 1 sh. ( ESFM-GAS/2014-53 ) ; 27.10.2014, Güllük Bay, Aegean Sea , sandy mud, 50 m, 4 sh. ( ESFM-GAS/2014-49 ) ; 07.02.2015, Ildır Bay, Aegean Sea , sandy mud, 66 m, 1 spm ( ESFM-GAS/2015-25 ) ; 08.02.2015, Ildır Bay, Aegean Sea , sandy mud, 67 m, 1 sh. ( ESFM-GAS/2015-31 ) ; 16.02.2015, G̹ll̹k Bay, Aegean Sea , sandy mud, 44 m, 2 spm + 1 sh. ( ESFM-GAS/2015- 33 ) ; 15.12.2015, Güllük Bay, Aegean Sea , sand, 45 m, 1 sh. ( ESFM-GAS/2015-32 ) ; 06.04.2017, Eski Foça, Aegean Sea , P. oceanica , 8 m, 1 spm ( ESFM-GAS/2017-87 ) ; 24.08.2017, G̹ll̹k Bay, Aegean Sea , sandy mud with shell fragments, 45 m, 1 spm ( ESFM-GAS/2017-88 ) ; 28.08.2019, G̹ll̹k Bay, Aegean Sea , sandy mud with shell fragments, 38 m, 3 spm ( ESFM-GAS/2019-18 ) ; 10.09.2019, Fethiye Bay , mud, 44 m, 1 spm ( ESFM-GAS/2019-21 ) .

Remarks: Among the Mangelia species distributed in the Mediterranean Sea, M. costulata is the easiest to identify due to its equal, thin and numerous spiral cords. The shell has a multispiral protoconch and orthocline axial ribs on teleoconch whorls (on the body whorl 9–10 ribs).

Distribution: Eastern Atlantic Ocean and Mediterranean Sea ( Gofas et al. 2011). Turkish coasts: Levantine Sea, Aegean Sea and Sea of Marmara (Özt̹rk et al. 2014).

Figures

FIGURE 6: Mangelia costulata: ventral (A, B, C, D,) views of four specimens and protoconch (E) of specimen D (A=6.6 mm, Nemrut Bay (Aliağa), 15 m; B= 5 mm, Iskenderun Bay, 11 m; C=4.7 mm, D=5.2 mm, Bodrum, 13 m).