Anamalysia mellipes van Achterberg & Yaakop, 2022

Yao, Junli, van Achterberg, Cornelis, Yaakop, Salmah, Long, Khuat Dang, Sharkey, Michael J. & Chapman, Eric G., 2022, A new genus Anamalysia van Achterberg (Hymenoptera, Braconidae, Alysiinae), six new species, and two new combinations from India, Indonesia, Malaysia, Singapore, Thailand, and Vietnam, ZooKeys 1126, pp. 131-154 : 131

Anamalysia mellipes van Achterberg & Yaakop sp. nov.

Fig. 3A-H

Type material.

Holotype, ♀ (RMNH), Malaysia, SW Sabah, near Long Pa Sia (West), c. 1050 m, 25.xi-8.xii.1987, Mal. trap 3, C. v. Achterberg, RMNH’ 87, DNA voucher number “94”. Paratypes (5 ♀): 1 ♀ (TAMU), Indonesia, West Kalimantan, Gunung Palung Nat. Park, 15.vi-15.viii.1991, Darling, Sutrisno & Rosichon, IIS 910122; Cabang Panti Res. Station, 1° [= primary] rainforest, 100-400 m, alluvial-light gap, 1°15'S, 110°5'E, Malaise trap, head; 1 ♀ (UKM), Malaysia, N. Sembilan, Pasoh Forest Reserve, 24.x.2002, (50 ha plot), Ng, Y.F. & Ruslan, DNA voucher number “84”; 1 ♀ (UKM), [West Malaysia], Johor: Endau-Rompin Selai, 25.ix-1.x.2004, Shah, Roslan, Fauzi, DNA voucher number “59”; 1 ♀ (RMNH), W. Malaysia, Johor, Endau Rompin, Kampung Peta, ix.2007, Mal. trap, Ruslan, Fauzi & Norlie; 1 ♀ (RMNH), W. Malaysia, Pahang, Hutan Kuala Lompat, 29.xii.2006-13.i.2007, Mal. Tr., Ruslan, Fauzi & Norlie, DNA voucher number “73”.

Description.

Holotype, ♀, length of body 3.6 mm, length of fore wing 3.1 mm.

Head. Width of head 1.7 times its median length, sparsely setose and strongly shiny; antenna incomplete, 24+, segments densely setose, length of third segment 0.9 times as long as fourth segment, length of third and fourth segments 4.0 and 4.3 times their width, respectively (Fig. 3E ); length of maxillary palp of paratypes 1.4 times height of head (missing in holotype); eye in dorsal view 4.4 times as long as temple; temple in dorsal view subparallel-sided behind eyes (Fig. 3B ); OOL: diameter of ocellus: POL = 11:4:3; frons flat medially (except a complete median groove) and convex laterally, smooth and no pit between antennal sockets; antennal sockets distinctly protruding; with a smooth, narrow and superficial groove between antennal sockets and eye; minimum width of face 0.5 times maximum width of head, densely rugulose-punctate submedially, more sparsely on remainder of face and transversely rugose ventrally and smooth medially, with rather long setae, without crenulate grooves ventrally; clypeus narrow, nearly parallel-sided, with long setae and ventrally rounded and its surface largely smooth except a few punctures (Fig. 3H ); length of malar space 0.1 times basal width of mandible; mandible rugose medially, strongly widened dorsally, its medial length 1.5 times its maximum width, upper tooth large and truncate lobe-shaped, with ventral tooth rather small, rounded and lobe-shaped, connected to a carina (Fig. 3C, G ).

Mesosoma. Length of mesosoma 1.5 times its height; pronotum dorsally with large deep and round dorsope; side of pronotum with some coarse crenulae anteriorly and medially, posteriorly finely crenulate and remainder smooth; epicnemial area dorsally smooth, medially crenulate and ventrally punctate-rugose; precoxal sulcus widely crenulate anteriorly, narrowed after its middle and absent posteriorly; remainder of mesopleuron smooth; episternal scrobe round, deep; pleural sulcus coarsely crenulate; mesosternal sulcus coarsely crenulate, rather wide posteriorly; metapleuron largely smooth, with some rugae ventrally; notauli complete, deep, narrow, and smooth; mesoscutum strongly shiny and largely glabrous, but with some long setae near notauli and lateral carina and medial lobe protuberant; mesoscutum without a separate medio-posterior depression; axilla rather setose and lateral carina moderately protuberant; scutellar sulcus deep, with one carina and no punctures, 0.4 times as long as scutellum; scutellum rather convex in lateral view; metanotum distinctly lamelliform protruding posteriorly in lateral view; surface of propodeum largely smooth anteriorly, except for a short median carina, medially with wide triangular areolate area and posteriorly reticulate; propodeal spiracle round, small and submedially at propodeum.

Wings. Pterostigma subelliptical (Fig. 3A ), its posterior margin hardly curved; vein r issued distinctly behind middle of pterostigma and distinctly oblique; r:3-SR:SR1 = 5:21:53; 1-SR+M rather sinuate; SR1 straight; cu-a short and oblique, interstitial; 2-SR:3-SR:r-m = 25:21:13; m- cu slightly postfurcal, slightly converging to 1-M posteriorly; first subdiscal cell 6.5 times as long as wide; CU1b distinctly shorter than 3-CU1 and 3-CU1 oblique. Hind wing: M+CU:1- M:1r-m = 30:35:13; m-cu distinctly developed and removed from 2-M.

Legs. Outer side of hind coxa largely smooth, punctulate and moderately setose, dorsally shiny and smooth; middle coxa strongly protruding forwards ventrally, hind coxa gradually narrowed; tarsal claws moderately robust (Fig. 3D ); length of femur, tibia, and basitarsus of hind leg 4.2, 14.0, and 10.3 times their width, respectively; hind tibia and basitarsus rather short and adpressed setose.

Metasoma. Length of first tergite 1.4 times its apical width, its surface regularly costate-striate, its dorsal carinae nearly complete and united submedially (Fig. 3F ); laterope absent; dorsope large and deep (Fig. 3F ); remainder of metasoma smooth and rather depressed; ovipositor with minute dorsal notch and some ventral teeth; length of setose part of ovipositor sheath 0.34 times fore wing and 0.8 times as long as hind tibia; apex of ovipositor sheath subtruncate and no apical spine; hypopygium medium-sized and apically acute in lateral view.

Colour. Dark chestnut brown; scapus, pedicellus, and tegulae more or less brown; palpi pale yellowish; metasoma (except first tergite and base of second tergite), remainder of antenna (as far as present), mandible, coxae (but paler apically) and ovipositor sheath brown; two basal segments of hind tarsus darkened; remainder of legs brownish yellow; pterostigma and most veins pale brown; wing membrane subhyaline.

Variation. Length of fore wing 2.8-3.1 mm and of body 3.3-3.6 mm; antenna of ♀ with 36 (1) segments, 1.9 times as long as fore wing and seven or eight apical segments white or ivory; vein SR1 of fore wing 2.5-3.1 times vein 3-SR; length of first tergite 1.4-1.5 times its apical width; eye in dorsal view 4.2-4.4 times as long as temple; length of setose part of ovipositor sheath 0.32-0.35 times fore wing.

Distribution.

Malaysia (East Malaysia: Sabah; West Malaysia: Johor, Pahang, Sembilan).

Etymology.

Named after its largely brownish yellow legs; "mel, mellis" is Latin for honey, and "pes, pedus" is Latin for leg.

Figures

Figure 3. Anamalysia mellipes sp. nov., ♀, holotype A fore wing B head, dorsal aspect C mandible, full sight on first tooth D outer hind claw E basal antennal segments F first metasomal tergite, dorsal aspect G mandible, full sight on third tooth H clypeus. Scale bars: 1.0 mm (A, B); 1.5 mm (C, E-G); 2.5 mm (D, H).