Heterocerus Fabricius, 1792

Sazhnev, Alexey S., 2020, Variegated mud-loving beetles (Heteroceridae) of Russia and adjacent countries Additions and corrections to the Catalogue of Palaearctic Coleoptera, Volume 3 (2016), Zootaxa 4810 (2), pp. 368-374 : 370-371

Genus Heterocerus Fabricius, 1792

p. 614: Heterocerus fausti Reitter, 1879 ; validity taxon, E: AB A: TM.

Distribution: Azerbaijan, Turkmenistan ( Sazhnev 2018c).

The validity of Heterocerus fausti Reitter, 1879 , previously considered a junior synonym of H. flexuosus Stephens, 1828 has been restored ( Sazhnev 2018c). The records of Heterocerus fausti for Uzbekistan ( Jacobson 1913; Zaitsev 1946) are not confirmed.

p. 613: Heterocerus fenestratus Thunberg, 1784 (Dermestes) ; add E: AB A: KI NTR

Distribution: Azerbaijan: Geok-tapa, Mingachevir, Jaffarkhan, Lankaran ( Zaitsev 1946); Karadag District Gobustal District ( Sazhnev 2018a); Kyrgyzstan: Issyk-Kul Oblast ( Ovchinnikov 1996), Jalal-Abad Oblast, Talas Oblast ( Litovkin & Sazhnev 2018); NTR: Chile (Sazhnev 2019).

p. 614: Heterocerus flexuosus Stephens, 1828 ; add E: NT A: KI.

Distribution: Russia: Republic of Karelia (Sazhnev 2018); Murmansk Oblast (Jakobson 1913); Kyrgyzstan: ( Ovchinnikov 1996); Issyk-Kul Oblast ( Litovkin & Sazhnev 2018).

p. 614: Heterocerus fossor Kiesenwetter, 1843 ; add E: BY UK.

Distribution: Belarus ( Alexandrovitch et al. 1996); Ukraine: Kiev, D-r. Lgocki (ZMMU); Nikolayev Oblast, Kinburn Spit, 29.06.2007 V. Dyadichko (IBIW).

Distributed in Europe, Turkey, Iran, Kazakhstan, Tajikistan, Siberia ( Mascagni, 2014, 2016). The record “Kirg.” for this species in the Jacobson’s Catalog (1913) refers to the Kirghiz steppe. It was recorded from Kyrgyzstan (North Kirghizia, Issyk-Kul hollow and Inner Tien-Shan) by Ovchinnikov (1996), but these records require verification ( Litovkin & Sazhnev 2018).

p. 614: Heterocerus fusculus fusculus Kiesenwetter, 1843 ; add E: NT A: KI TD TM UZ.

Distribution: Russia: Leningrad Oblast ( Zaitsev 1907); St.- Petersburg, Lesnoy, 20.05.1862, А. Moravits (ZISP), and 29.05.1921, A. Vulf (ZISP); Kyrgyzstan: (Zaitsev 1910; Ovchinnikov 1996); Issyk-Kul Oblast ( Litovkin & Sazhnev 2018); Tajikistan: Dusti, 29– 30.08.1989, Kononenko (SZM); Turkmenistan: Repetek, 05.05.1971, А. Kurentsov (MSPU); Uzbekistan ( Prokin et al. 2016).

p. 614: Heterocerus heydeni Kuwert, 1890 ; validity taxon, E: ST KZ UK (Crimea) A: KZ TM UZ.

New taxonomic data on mud-loving beetles are provided based on morphological characters and DNA barcoding. Heterocerus heydeni was previously considered a junior synonym of Heterocerus flexuosus , but the authors support the validity of the species and restore the name. Specimens of Heterocerus hauseri Kuwert, 1893 were also studied, suggesting it as a possible junior synonym of Heterocerus heydeni ( Litovkin et al. 2019) .

p. 614: Heterocerus kamtschaticus Egorov, 1989 ; a junior synonym of Heterocerus fenestratus Thunberg, 1784 ( Sazhnev 2016b) .

p. 615: Heterocerus kaszabi R. Charpentier, 1979 ; add A: WS.

Distribution: Russia: Republik of Buryatia, Beloozersk, Beloe Lake, 1– 7.07.1999, I. Melnik (MSPU); Zabaykalsky Krai (Sazhnev 2016).

p. 615: Heterocerus marginatus Fabricius, 1787 (Apate) ; add E: BY NT A: FE MG.

Distribution: Belarus ( Alexandrovitch et al. 1996); Russia: Arkhangelsk Oblast ( Pomerantsev 1908); Leningrad Oblast ( Lindemann 1871; Zaitsev 1907); Vologda Oblast (Sazhnev 2018); Ussuriysk, Kamenushka, 07.07.1980, A. Ryvkin (IBIW); Mongolia: Dornod Province ( Prokin et al. 2019).

p. 615: Heterocerus mus Charpentier, 1965 ; add A: KI. Distribution: Kyrgyzstan: Jalal-Abad Oblast ( Litovkin & Sazhnev 2018).

p. 615: Heterocerus nepalensis A. Mascagni, 1993 ; add A: KI. Distribution: Kyrgyzstan ( Skalický 2010).

p. 615: Heterocerus obsoletus Curtis, 1828 ; add E: AR NT A: KI UZ.

Distribution: Armenia: Tsakhkadzor mountain range, Teckyarlu, 22– 23.06.1997, I. Melnik (MSPU); Russia: Republic of Karelia (Sazhnev 2018); Murmansk Oblast (Sazhnev 2018); Kyrgyzstan: ( Ovchinnikov 1996); Issyk- Kul Oblast, Jalal-Abad Oblast ( Litovkin & Sazhnev 2018); Uzbekistan ( Prokin et al. 2016).

p. 615: Heterocerus parallelus Gebler, 1830 ; add E: UK A: KI TR.

Distribution: Ukraine: Kherson Oblast, Dzharylhach Island, 18.08.2002, V. Mikhailov (IBIW); Rybachie, 17.05.– 15.06.2003 M. Makarenko (MSPU); Donetsk Oblast, Samsonovo, 17.08.1990 D. Novokhatsky (MSPU); Kyrgyzstan: ( Ovchinnikov 1996); Issyk-Kul Oblast ( Litovkin & Sazhnev 2018); Turkey: Gyumushkhana ( Zaitsev, 1946).

add Heterocerus virgatus Mamitza, 1933 ; A: KI.

Distribution: Kyrgyzstan ( Skalický 2010).