Pterostichus (Badistrinus) goschi

Makarov, K. V. & Sundukov, Yu. N., 2022, On status and systematic position of Pterostichus goschi Jedlička, 1930 (Coleoptera: Carabidae), Russian Entomological Journal 31 (4), pp. 362-372 : 363-372

Pterostichus (Badistrinus) goschi

Jedlička, 1930, stat.resurr.

Figs 1–18 , 25–30, 33–35 , 45–50 .

Pterostichus ( Badistrinus ?) Goschi Jedlička, 1930: 21 ; type locality: “Sibiria or.: Nikolsk-Ussurijsk”, synonymy established for Pterostichus neglectus A. Morawitz, 1862 by Sundukov, 2013: 64, stat.resurr.

Pterostichus (Badistrinus) bandotaro Tanaka, 1958: 78–79 , fig. 1; type locality: “ Lake Wadanuma in the river-bed of the Toné, Chiba Pref., Honshu, Japan ”, syn.n .

Pterostichus (Phonias) neglectoides Budilov, 2022: 25–27 , figs 1–9; type locality: “ Russia: Jewish Autonomous Region, the protected zone of´Bastak´ Reserve”; synonymy established for Pterostichus bandotaro Tanaka, 1958 by Sasakawa, 2022: 19.

TYPE MATERIAL. Lectotype (designated herewith), ♂, with the following labels: “Nikolsk Ussurijsk Ussurigeb. Mandl” [white rectangle; printed], “ goschi mihi, DET. ING. JEDLIČKA” [pink rectangle; handwritten/printed], “TYPE” [brown rectangle with a black border; printed], “Mus. Nat. Pragae Inv. 22370” [yellow rectangle; printed/handwritten], “ LECTOTYPUS Pterostichus (Badistrinus) goschi Jedlička, 1930 design. K. Makarov et Yu. Sundukov, 2022” [red rectangle with a black border; printed] ( NMPC). Paralectotype, ♂, with the following labels: “Nikolsk Ussurijsk Ussurigeb. Mandl” [white rectangle; printed], “ goschi mihi, det. Ing. Jedlička” [white rectangle; handwritten/printed], “TYPE” [brown rectangle with a black border; printed], “Mus. Nat. Pragae Inv.22369” [yellow rectangle; printed/handwritten], “ PARA- LECTOTYPUS Pterostichus (Badistrinus) goschi Jedlička, 1930 design. K. Makarov et Yu. Sundukov, 2022” [red rectangle with a black border; printed] ( NMPC).

OTHER MATERIAL. Russia: Primorsky Krai: Khasansky district, Khasan , at light, 25.VII.1970, Yu.N. Nazarov leg., 3♂♂ 7♀♀ ( ZISP) ; ibid, Golubiny Utes, 2–2.5 km on the way to Khasan , 24.V.2022, Yu. Sundukov leg., 3 ex ( FEB) ; ibid, Golubiny Utes , traps in meadow, 22–24.V.2022, Yu. Sundukov leg., 3 ex ( FEB) ; ibid, Golubiny Utes , at night on the coast, 22.V.2022, Yu. Sundukov leg., 2 ex ( FEB) ; ibid, 10.7 km east of Khasan, Cape Ostrovok Falshivy , 42°26´54´´N 130°46´47´´E, 23.V.2022, K. Makarov leg., 1 ex ( MSPU) ; ibid,1.7kmnorthofCapeL’va, 42°42´15´´N 131°14´12´´E, 28.V.2022, K. Makarov leg., 1 ex ( MSPU) ; ibid, near Zanadvorovka, Kedrovaya River valley , 43°18´53´´N 131°34´43´´E, 9–17.VII.2000, I. Melnik leg., 1 ex ( MSPU) ; Nadezhdensky district, lower reaches of Malaya Klyopochnaya River near Razdolnoe , 5.IX.2019, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; “near Vladivostok , 25.IV.1911, A. Chersky leg.”, 1♂ ( ZISP) ; “ Vladivostok ”, 2♂♂, 1♀ ( ZISP) ; “ Vladivostok 22 III 70”, 3♂♂, 3♀♀ ( ZISP) ; “ Vladivostok 15.V.70”, 1♀ ( ZISP) ; “ Odarkovsky zavod, Prim. ob., 3.VII.1911, A. Chersky leg.”, 1♂ ( ZISP) ; “ Nikolsk Ussur. A. Ivanov 5.VIII.26”, 1♀ ( ZISP) ; “okr. Chernigovki Prim. o. Emel´yanov 20.VI.912”, 1♂ ( ZISP) ; “ Troitskoe, oz. Khanka, Uss. kr. Chersky 5.IX.914”, 1♂ ( ZISP) ; near Beltsovo, lower reaches of Arsenievka River , right bank, floodplain, 28.VII.2003, Yu. Sundukov & V. Shokhrin leg., 3 ex ( FEB) ; Lazovsky district, near Lazo , 6–10.VI.2008, V. Shokhrin leg., 1 ex ( FEB) ; Khabarovsky Krai : “ Khabarovsk, Primorsk. obl. 7.VI.11 Speshilova-Petelina”, 1♀ ( ZISP) ; Komsomolsky Reserve, cordon Bichi, bank of Gorin River , traps, 25–30.VII.2021, V. Bobrovsky leg., 3 ex ( KSR) ; ibid,a stream above the 3rd Byk on Gorin River , 26.VII.2021, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; ibid, unnamed stream at the 2nd Byk , 28.VII.2021, O. Kuberskaya leg., 2 ex ( KSR) ; ibid,a stream between the 1st and 2nd Byk on Gorin River , 28.VII.2021, Yu. Sundukov & L. Sundukova leg., 4 ex ( FEB) ; ibid, Khawar Stream below cordon Bichi , 24.VII.2021, Yu. Sundukov & L. Sundukova leg., 2 ex ( FEB) ; ibid, 24.VII.2021, O. Kuberskaya leg., 3 ex ( KSR) ; ibid, Namek Stream , 23.VII.2021, O. Kuberskaya leg., 4 ex ( FEB, KSR) ; ibid, Chenki, bank of Amur River , 29.VII.2021, O. Kuberskaya leg., 10 ex ( FEB, KSR) ; ibid, Strelka , willow, 24.VII.2021, O. Kuberskaya leg., 2 ex ( KSR) ; ibid, cordon Tikhaya , 2.VIII.2021, O. Kuberskaya leg.,1 ex ( KSR) ; ibid, Delfin ,wasteland near the children´s camp, 2–14.VII.2021, O. Kuberskaya leg., 2 ex ( KSR) ; Komsomolsk-na-Amure, Silinsky Park, willow sandy bank of Silinka River , 6– 13.VII.2021, O. Kuberskaya leg., 1 ex ( KSR) ; Southern Kuriles, Kunashir Island: Lovtsova Peninsula , a lake north of Lake Nadya , 8.VIII.2015, Yu. Sundukov & L. Sundukova leg., 4 ex ( FEB) ; ibid, swamp in southern part of Lake Nadya , 15–16.VIII.2015, Yu. Sundukov & L. Sundukova leg., 16 ex ( FEB) ; ibid, NE coast of Lake Dlinnoe , meadow, 18.VIII.2015, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; forest lake 1 km above the mouth of Tyatina River , 08.VI.2016, Yu.Sundukov leg., 13 ex ( FEB) ; Tret'yakolvo , 5.08.1973, I.M. Kerzner, 1 ♀ ( ZISP) ; lower lake in stream valley 2 km south of western bank of Lake Peschanoe , 3.VII.2015, Yu. Sundukov & L. Sundukova leg., 9 ex ( FEB) ; upper and middle lakes in stream valley 2 km south of the western bank of Lake Peschanoe , 4.VII.2015, Yu. Sundukov & L. Sundukova leg., 45 ex ( FEB) ; middle lake in stream valley 2 km south of western bank of Lake Peschanoe , 26.VII.2015, Yu. Sundukov leg., 9 ex ( FEB) . Japan: “ Hakodate 110-3.”, “ neglectus Mor. typ. A. Morawitz det.”, “ Pterostichus bandotaro Tanaka, Det. S. Morta, 2012 ”, 1♂ ( ZISP) .

Comparative material on Pterostichus neglectus A. Morawitz, 1862 .

TYPE MATERIAL. Lectotype (designated herewith), ♂, with the following labels: “M. Bureja 110-1.” [red rectangle in a black frame; handwritten], “ neglectus Mor. typ. A. Morawitz det.” [white rectangle; handwritten], “ Pterostichus neglectus Morawitz, Det. S. Morita, 2012 ” [white rectangle; printed], “ LECTOTYPUS Pterostichus (Badistrinus) neglectus A. Morawitz, 1862 design. K. Makarov et Yu. Sundukov, 2022” [red rectangle with a black border; print- ed]” ( ZISP) ( Figs 19–20 ). In the original description [ Morawitz, 1862], two type localities are indicated — “Bureya” and the mouth of the Amur River. In addition, from the text of the description (“Der abgekürtze Scutellanstreif fehlt bei einem Exemplare, und ist bei den andern nur als kurzes, schrägers Stichelchen dicht am zweiten Streifen forhanden”) it is clear that several specimens were at Moravitz´s disposal, but their exact number is not indicated. So, we designate a specimen from the ZISP collection, the label of which corresponds to the first mentioned type locality, as a lectotype.

OTHER MATERIAL. Russia: Primorsky Krai: Khasansky district, 18 km SW Barabash, the floodplain of the Bezymyanny Stream , a swamp near the highway, 16.VIII.2020, Yu. Sundukov leg., 1 ex ( FEB) ; ibid, 43°3´52´´N 131°21´49´´E, 16.VIII.2020, K. Makarov leg., 1 ex ( MSPU) ; ibid, cordon Siny Utes , sedge swamp in forest, 26.V.2021, Yu. Sundukov leg., 1 ex ( FEB) ; Borisov Plateau, watershed of the Valunny Ridge opposite mouth of Kamenisty Stream , h~ 600–650 m, 28.V.2019, Yu. Sundukov leg., 2 ex ( FEB) ; Lazovsky district, near Lazo , 28–29.VI.2006, Yu. Sundukov leg., 2 ex ( FEB) ; ibid, 6–10.VI.2008, V. Shokhrin leg., 1 ex ( FEB) ; Spassk-Dalnii , at light, 10.VII.2016, M. Sergeev leg., 1 ex ( FEB) ; Sikhote-Alinsky Reserve, shore of Lake Tsarskoye , 9.X.1982, A. Turchinskaya leg., 1 ex ( MSPU) ; ibid, near Terney, cordon Maysa , 10.X.1982, A. Matalin leg., 2 ex ( MSPU) ; Khabarovsky Krai: Khabarovsk, Industrialnaya str., on muddy shore, end of June 1997, E. Novomodny leg., 1 ex ( KTM) ; Komsomolsk-na-Amure, Silinsky Forest , broad-leaved forest near southern border, 27.IV– 5.V.2020, O. Kuberskaya leg., 1 ex ( KSR) ; ibid, Silinsky Park , lake bank, 22.VII.2021, Yu. Sundukov leg., 1 ex ( FEB) ; Komsomolsky Reserve, Khawar Stream below cordon Bichi , 24.VII.2021, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; ibid, 24.VII.2021, O. Kuberskaya leg., 1 ex ( KSR) ; ibid, Shargol channel near Delfin base, 22.VII.2021, O. Kuberskaya leg., 1 ex ( KSR) ; ibid, cordon Tikhaya , 2.VIII.2021, O. Kuberskaya leg., 1 ex ( KSR) ; ibid, Chenki, on Amur River , 29.VII.2021, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; ibid, mouth of the Gorin River ( Strelka ), willow, 25.VII.2021, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; ibid, a stream between the 1st and 2nd Byk on Gorin River , 28.VII.2021, Yu. Sundukov & L. Sundukova leg., 1 ex ( FEB) ; ibid, cordon Bichi, bank of Gorin River , 23.VII.2021, Yu. Sundukov & L. Sundukova leg., 2 ex ( FEB) ; Amur Oblast: Khingansky Reserve, Khingan forestry, Karabcha River valley , reed grass with herbs, traps, 13– 23.VI.2000, E. Ignatenko leg., 2 ex ( FEB) ; Zabaykalsky Krai: Chita area, near Duroy , 50°00´30´´N 118°55´37´´E, 16–18.VI.2000, I. Melnik leg., 1♂ ( MSPU) ; ibid, Priargunsky district, Duroy , 16– 18.VI.2000, B. Kataev, 1♂ ( ZISP) .

LECTOTYPE REDESCRIPTION ( Figs 1–11 ). The body moderately convex. Length 5.2 mm, width 2.0 mm.

Body black, shiny; antennae darkened, with reddish-brown basal segment and segments 2–4 proximally; palps brown, apical segment of maxillary palps bicolored: with darkened base and lightened apex; mandibles blackish, with brown apices and outer margin; upper lip brownish black. Underside black. Legs bicolored: coxae, trochanters and femora blackish-brown, tibiae and tarsi reddish-brown.

Dorsum of body with very fine scattered punctures: dens- er on head and in the disk of pronotum, strongly scattered on sides of pronotum and elytra; basal pits of pronotum and area between them, with large, rather sparse punctures. Microsculpture on upper side of head and pronotum practically absent, weak traces of isodiametric reticulation visible only in depressions; elytra with very weak, strongly transverse microsculpture.

Dimensions (in mm): HW = 0.98; HL = 0.65; ML = 0.40; PA = 0.98; PW = 1.60; PB = 1.23; PL = 1.28; EW = 1.98; EL = 3.03; L = 5.23.

Head small, moderately convex, its width with eyes 1.51 times its length. Eyes large, strongly protruding. Mandibles small, with pointed apices bent inwards. Labrum slightly trapezoid; its anterior margin uniformly weakly convex, with 6 setae. Clypeus trapezoid, with 2 lateral setae; its anterior margin slightly concave. Mentum median tooth short, bifid, with 2 setae at base. Submentum with 2 pairs of large setae, outer setae shorter. Gula in the center with thin, longitudinal groove. Temples very short, about 1/6 eye diameter. Supraorbital setae 2: 1/3 from anterior margin of eye and near its posterior margin. Frontal grooves weak, wide, not reaching anterior supraorbital seta. Antennae rather thin, reaching 1/5 of elytral length.

Pronotum subquadrate, with uniformly convex disc starting from narrow lateral margin, slightly transverse (PW/PL = 1.25), much wider than head (PW/HW = 1.63), with maximum width at the middle. Anterior margin very slightly concave, finely edged at anterior angles. Anterior angles slightly projecting forward, their apices widely rounded. Base rectilinear, edged near basal pits, wider than anterior margin (PB/PA = 1.26). Posterior angles obtuse, not protruding, widely rounded at apices. Lateral sides moderately evenly convex from anterior to posterior angles. Lateral margins narrowly evenly bordered. Lateral sides with 2 setae each: at 2/3 from the base and in posterior angle. Median line thin, deep, extending to anterior and posterior transverse depressions. Transverse depressions weak, indistinct. Basal pits represented by narrow, deeply depressed strokes; inner strokes 2.5 times longer than outer ones.

Elytra long, narrow (EL/EW = 1.53, EL/PL = 2.37, EW/PW = 1.24), almost parallel-sided, moderately convex, with a maximum width at 3/5 from base. Shoulders straight, without teeth. Basal edging complete, slightly concave. Lateral flattened margin narrow, slightly widening towards apex. Outer apical angle broadly rounded. Striae complete, narrow, deep, not punctate. Intervals convex, outer ones slightly explanate. Scutellar stria distinct, rather long, located in 2nd interval. Parascutellar pores located at the apex of scutellar stria, close to 2nd stria. 3rd interval with 2 small discal setigerous punctures located near 3rd stria: anterior one approximately at the middle of elytron, posterior one approximately at 1/5 from apex. Apex of stria 7 with one preapical and one apical setigerous punctures. Lateral series (series umbilicata) consisting of 13 setae: 5+ 1 in humeral group and 3+4 at apex. Full-winged.

Legs moderately long, rather slender; length of hind tarsus approximately equal to length of hind tibia.

Underside of body not punctured. Anterior metasternal process not edged, widely rounded at apex. Metepisternae long, narrow: 2.7 times as long along outer margin as wide along anterior margin. Hind coxae with 2 setae. Posterior trochanters with 1 seta. Abdominal sternites simple, without pubescence and additional setae; apical sternite with 2 setae at apex.

Aedeagus ( Figs 4–8 ) rather slender, curved at right angle; lamella symmetrical, short, distinctly narrowed towards rounded apex. Left paramere wide, discoid; right paramere narrow.

Endophallus ( Figs 9–11 ) narrow, elongate, oriented almost at right angle to the plane of lamella and prepuce. Basal part cracked during preparation, but its shape was not distort- ed. Medial part with paired lateral protuberances, slightly asymmetrical, below them endophallus clearly narrowed. Distally, endophallus slightly expanded, bearing a complex of strongly protruding tubercles. Ventro-apical protrusion (D) long, S-shaped, covered with sclerotized denticles and noticeably asymmetric, with small conical protrusion at the base on the left. Distally, endophallus surrounded by three asymmetric tubercles with weak sculpture: large left (C), small rounded mediodorsal (A), and strongly curved right (B). Towards gonopore, endophallus tube retaining asymmetry, distinctly inclined to the right, and bearing several tiers of large spines.

VARIABILITY.The paralectotype ( Figs 12–18 ) is slightly larger than the holotype (length 5.5 mm, width 2.1 mm). It has fairly distinct isodiametric reticulation on the head ( Fig. 12 ), barely visible, slightly transverse on the pronotal disc ( Fig. 13 ) and distinct transverse microsculpture on the elytra ( Fig. 14 ). Both third intervals have only 1 discal pore located near the middle of the elytron.

DISTRIBUTION. Russia: Jewish Autonomous Region, south of Khabarovsky Krai, Primorsky Krai, South Kuriles: Kunashir (new record); North Korea; Japan: Hokkaido, Honshu (Map). In connection with the separation of P. neglectus and P. goschi , records of P. neglectus from the south of Sakhalin require re-identification.

TAXONOMIC REMARKS. In the Far East, P. goschi lives sympatrically with P. neglectus A. Morawitz, 1862 (Map). The study of extensive collection material without dissection of the endophallus led to the erroneous conclusion that, in terms of the shape of the elytra, length of scutellar stria, and the coloration of legs and palps, individuals of P. goschi fit well into P. neglectus variability, which led to the reduction of P. goschi to junior synonyms of the latter species [ Sundukov, 2013].

The differences between P. neglectus and P. goschi were rightly pointed out by Budilov [2021]. However, he designat- ed them as “forms” of P. neglectus , and subsequently described one of them as an independent species [ Budilov, 2022]. After publications of Budilov [2022] and Sasakawa [2022], the authors again became interested in the synonymy of these taxa. For this purpose, syntypes of P. goschi were studied, in one of which it was possible to evert and study the endophallus ( Figs 9–11 ). Comparison of descriptions [ Budilov, 2022; Jedlička, 1930; Morawitz, 1862; Sasakawa, 2022; Tanaka, 1958] and endophallus structure ( Figs 21–28 ) confirmed both the differences between P. neglectus and P. goschi , and the conspecificity of the latter with P. bandotaro Tanaka, 1958 and P. neglectoides Budilov, 2022 . The following characters can be used to distinguish between these species:

1. Lateral sides of pronotum evenly rounded to posterior angles ( Figs 45–50 ), apices of posterior angles widely rounded ( Fig. 30 ); femora and partially palps darkened ( Fig. 29 ), trochanters and coxae entirely black; scutellar stria usually normally developed; concave side of the aedeagus more or less even ( Figs 33–35 ); endophallus ( Figs 9–11 , 25–28 ) ......................................... P. goschi

— Lateral sides of pronotum shortly concave or rectilinear in front of posterior angles ( Figs 39–44 ), apices of posterior angles shaped like teeth ( Fig. 32 ); legs and palps uniformly red ( Fig. 31 ), trochanters and coxae partially reddish; scutellar stria usually shortened, often to a single point; concave side of aedeagus swollen medially ( Figs 36–38 ); endophallus ( Figs 21–24 ) .......................... P. neglectus

It should be noted that these closely related species are currently included in two subgenera, Phonias Gozis, 1886 and Badistrinus Motschulsky, 1866 [ Bousquet, 2017]. The similarity of the endophallus structure plans (lack of a bend, large asymmetric tubercles, rows of spines in the apical part)

48 49 50

and the position of the apical discal pore (close to the 3rd stria) indicate that these species belong to the same subgenus and, at the same time, separate them from Phonias. At the present level of knowledge of the subgenus structure of Pterostichus Bonelli, 1810 , it seems appropriate to place these species in the subgenus Badistrinus .

Acknowledgements. We are sincerely grateful to Jiří Hájek (NMPC, Prague, Czech) and Boris Kataev (ZISP, St. Petersburg, Russia) for the possibility to study the types of P. neglectus and P. goschi . The research of Yu. Sundukov was carried out within the state assignment of Ministry of Science and Higher Education of the Russian Federation (theme No. 121031000151-3).

Competing interests. The authors declare no competing interests.

Figures

Figs 1–3. Pterostichus goschi Jedlička, 1930, lectotype, male: 1 — dorsal habitus; 2 — ventral habitus; 3 — labels. Рис. 1–3. Pterostichus goschi Jedlička, 1930, лектотип, самец: 1 — сверху; 2 — сниЗу; 3 — Этикетки.

Figs 4–8. Pterostichus goschi Jedlička, 1930, lectotype, male genitalia: 4 — aedeagus, left view; 5 — same, right view; 6 — same, dorsal view; 7 — right paramere; 8 — left paramere. Рис.4–8. Pterostichus goschi Jedlička, 1930, лектотип, гениталии самца: 4 — Эдеагус, слева; 5 — то Же, справа; 6 — то Же, сверху; 7 — праваЯ парамера; 8 — леваЯ парамера.

Figs 9–11. Pterostichus goschi Jedlička, 1930, lectotype, male endophallus: 9 — left lateral view; 10 — ventral view; 11 — right view. Designations for the endophallus structures (A–D) according to Budilov [2022]. Рис. 9–11. Pterostichus goschi Jedlička, 1930, лектотип, самец, Эндофаллус: 9 — слева; 10 — сниЗу; 11 — справа. ОбоЗначениЯ структур (A–D) по Budilov [2022].

Figs 12–18. Pterostichus goschi Jedlička, 1930, paralectotype, male: 12–17 — microsculpture (12 — forehead; 13 — disk of pronotum; 14 — 3rd elytral interval, 15–17 — first, second and third visible sternite); 18 — labels. Рис. 12–18. Pterostichus goschi Jedlička, 1930, паралектотип, самец: 12–17 — микроскульптура (12 — лоб, 13 — диск переднеспинки; 14 — 3й промеЖуток надкрылий; 15–17 — первый, второй и третий видимые стерниты); 18 — Этикетки.

Figs 21–28. Endophallus: 21–24 — Pterostichus neglectus A. Morawitz, 1862 (Khabarovsky Krai, Khawar Stream below cordon Bichi); 25–28 — P. goschi Jedlička, 1930 (25 — Nikolsk Ussurijsk, lectotype; 26 — Kunashir, forest lake 1 km above mouth of Tyatina River; 27 — Primorsky Krai, Cape Ostrovok Falshivy; 28 — Khabarovsky Krai, Namek Stream); 21 — left view; 22 — dorsal view; 23, 25–28 — ventral view; 24 — right view. Рис. 21–28. Эндофаллус: 21–24 — Pterostichus neglectus A. Morawitz, 1862 (Хабаровский край, ручей Хавар ниЖе кордона Бичи); 25–28 — P. goschi Jedlička, 1930 (25 — Уссурийск [Nikolsk Ussurijsk], лектотип; 26 — Кунашир, лесное оЗеро 1 км выше устьЯ р. ТЯтина; 27 — Приморский край, мыс Островок Фальшивый; 28 — Хабаровский край, ручей Намёк); 21 — слева; 22 — сверху; 23, 25–28 — сниЗу; 24 — справа.

Figs 29–38. Male of Pterostichus (Badistrinus) from Khasan district, detals: 29–30, 33–35 — P. goschi Jedlička, 1930; 31–32, 36– 38 — P. neglectus A. Morawitz, 1862; 29, 31 — dorsal habitus; 30, 32 — hind angle of pronotum; 33–38 — aedeagus, lateral and dorsal view. Рис. 29–38. Детали строениЯ самцов of Pterostichus (Badistrinus) иЗ Хасанского района: 29–30, 33–35 — P. goschi Jedlička, 1930; 31–32, 36–38 — P. neglectus A. Morawitz, 1862; 29, 31 — внешний вид, сверху; 30, 32 — Задние углы переднеспинки; 33–38 — Эдеагус, сбоку и сверху.

Figs 39–50. Variation in the shape of the pronotum: 39–44 – Pterostichus neglectus A. Morawitz, 1862 (39 – Primorsky Krai, 18 km SW Barabash; 40 – Khabarovsky Krai, Khawar Stream below cordon Bichi; 41 – Primorsky Krai, Sikhote-Alinsky Reserve, bank of Lake Tsarskoye; 42 – Zabaykalsky Krai, Priargunsky district, Duroy; 43 – Khabarovsky Krai, stream between the 1st and 2nd Byk on Gorin River; 44 – Zabaykalsky Krai, Chita area, near Duroy); 45–50 – P. goschi Jedlička, 1930 (45 – Primorsky Krai, Cape Ostrovok Falshivy; 46 – Khabarovsky Krai, Namek Stream; 47 – Kunashir Island, forest lake 1 km above mouth of Tyatina River; 48 – Primorsky Krai, 1.7 km north of Cape L’va; 49, 50 – Primorsky Krai, Khasan environs). Рис. 39–50. ИЗменчивость формы переднеспинки: 39–44 — Pterostichus neglectus A. Morawitz, 1862 (39 — Приморский край, 18 км ЮЗ Барабаш; 40 — Хабаровский край, руч. Хавар ниЖе кордона Бичи; 41 — Приморский край, Сихоте-Алинский Заповедник, берег оЗ. Царское; 42 — Забайкальский край, Приаргунский р-н, Дурой; 43 — Хабаровский край, ручей меЖду 1-м и 2-м Быками на р. Горин; 44 — Забайкальский край, окрестности п. Дурой); 45–50 — P. goschi Jedlička, 1930 (45 — Приморский край, мыс Островок Фальшивый; 46 — Хабаровский край, ручей Намёк; 47 — Кунашир, лесное оЗеро 1 км выше устьЯ р. ТЯтина; 48 — Приморский край, 1,7 км севернее мыса Льва; 49, 50 — Приморский край, окрестности п. Хасан).

Figs 19–20. Pterostichus neglectus A. Morawitz, 1862, lectotype, male: 19 — dorsal habitus; 20 — labels. Рис. 19–20. Pterostichus neglectus A. Morawitz, 1862, лектотип, самец: 19 — вид сверху; 20 — Этикетки.