Praearchitipula mirabilis, Gao, Jiaqi, Shih, Chungkun, Kopeć, Katarzyna, Krzemiński, Wiesław & Ren, Dong, 2015

Gao, Jiaqi, Shih, Chungkun, Kopeć, Katarzyna, Krzemiński, Wiesław & Ren, Dong, 2015, New species and revisions of Pediciidae (Diptera) from the Middle Jurassic of northeastern China and Russia, Zootaxa 3963 (2), pp. 240-249 : 247

Praearchitipula mirabilis sp. nov.

( Fig. 5 )

Diagnosis. Cross-vein sc-r at a short distance distal from the origin of Rs; cross-vein r-m reaching R5 not far from the forking of R4+5; cell m1 slightly longer than petiole of cell m1 (dM1+2); M1+2 only 1/3 as long as M1; d-cell about 1/ 7 of the length of wing; cross-vein m-cu just proximad of the bifurcation of M3+4. (Note: The positions of sc-r and m-cu are unique features distinguishing this new species from the other species of this genus.)

Etymology. The specific name is derived from a Latin word “ mirabilis ”, referring to the peculiar and different position of cross-vein sc-r distal of the origin of Rs.

Description. A medium-sized crane fly, body length 8 mm with well-preserved wings. Head oviform, but features of the head (e.g. tip of the rostrum, palpi and antennae) unidentifiable; compound eyes preserved. The thorax well-preserved; the boundaries between prothorax, mesothorax and metathorax distinct in lateral view; mesonotum robust and well-developed; some parts of mesopleura distinguishable; halter spoon-shaped, 1.4 mm long. The legs long and slender. The abdomen poorly preserved, relatively long and slender, with 8 clear segments. Female with ovipositor discernible. Wing: The right wing well-preserved. Length 6.7 mm and width 2.4 mm, 2.8 times as long as wide. Wing clear with light-colored pterostigmal region. Venation: Sc long, 0.71 times of the wing length and ending distad of the level of branching of R4+5; cross-vein sc-r far from the tip of Sc, at a short distance distal from the origination of Rs; Rs of moderate length, strongly arcuated at origin; Rs arising from 0.33 times the length of wing, Rs bifurcates into R2+3 and R4+5 at 0.66 times length of wing,; R1 long; r-r (R2) close to R1 tip, transverse; R4+5 short, 0.19 times as long as R2+3. Stem of vein M bifurcating at the same level of the branching of Rs; cell m1 petiolate; mM1+2 nearly as long as cell m1, cell m1 1.2 times as long as petiole of cell m1 (dM1+2); crossvein r-m entering R5 not far from the forking of R4+5, dR5 twenty times as long as bR5; discal cell long, about 1/7 of the wing length; cross-vein m-cu distinctly proximad of M3+4 bifurcation, bM3+4 4 times as long as dM3+4; cross-vein m-cu close to the bifurcation of M3+4; Cu long; anal veins long and divergent.

Material examined. Holotype, female, CNU-DIP-NN2014003 ( Fig. 5 ), a partly preserved specimen of a hairy-eyed crane fly in lateral view, with partly-preserved abdomen, the wing clearly visible as preserved.

Figures

FIGURE 5. Praearchitipula mirabilis sp. nov., holotype, CNU-DIP-NN 2014003. A—Photograph; B—Enlarged details (sc-r) of right wing; C—Enlarged details (sc-r) of left wing; D—Line drawing of habitus; E—Line drawing of right wing. Scale bars = 1 mm.