Himaloconnus klapperichianus amamianus, Jałoszyński, 2020

Jałoszyński, Paweł, 2020, Himaloconnus Franz and Nogunius gen. n. of Japan (Coleoptera: Staphylinidae Scydmaeninae), Zootaxa 4822 (3), pp. 334-360 : 339-342

Himaloconnus klapperichianus amamianus ssp. n.

( Figs 2 , 6–7 , 36–39)

Type material. Holotype: JAPAN (Kagoshima Pref.): ♂, two labels: “JAPAN, KAGOSHIMA Pref. / Amami-Ôshima, Yui-dake / 25-28.v.2004, white pan trap / Toshiharu Mita leg.” [white, printed], “ HIMALOCONNUS / klapperichianus amamianus m. / HOLOTYPUS, P. JAŁOSZYŃSKI, 2020” [red, printed] ( NSMT) . Paratypes (5 exx.): 2 ♂♂, same data as for holotype; 3 ♂♂ (one as whole-body mount in Canada balsam), “JAPAN, KAGOSHIMA Pref. / Amami-Ôshima, Kinsakubaru, / Naze-shi, yellow pan trap / 29.v.-1.vi.2004 / Toshiharu Mita leg.” ( NSMT, cPJ).

Diagnosis. BL of male <1 mm; eyes in males in dorsal view slightly longer than length of tempora, each composed of conspicuously large ommatidia ( Fig. 7 ).

Description. Body of male ( Fig. 2 ) strongly convex, uniformly light brown, covered with setae distinctly lighter than cuticle; BL 0.88–0.96 mm (mean 0.90 mm).

Head ( Figs 6–7 ) broadest at eyes, HL 0.16–0.19 mm (mean 0.18 mm), HW 0.18–0.19 mm (mean 0.18 mm); tempora in dorsal view slightly shorter than eyes; vertex and frons confluent, together weakly convex; each eye ( Fig. 7 ) large, strongly convex, bean-shaped, with distinct posteromedian emargination, composed of 23 conspicuously large facets. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; tempora and posterior margin of vertex densely covered with long and thick bristles. Antennae slender, AnL 0.38–0.40 mm (mean 0.39 mm), antennomeres I and II each strongly elongate, III–VIII each indistinctly transverse, IX and X distinctly transverse, XI indistinctly narrower than X, slightly elongate.

Pronotum subconical with rounded sides, broadest near posterior third, nearly evenly tapering anterad; PL 0.25 mm, PW 0.23 mm. Anterior margin nearly straight, posterior margin indistinctly bisinuate with weakly convex median third, base lacking pits. Punctures on pronotal disc fine, inconspicuous; setae sparse, moderately long, suberect; sides with sparse, thick and long bristles.

Elytra together oval, broadest at or near middle; EL 0.45–0.53 mm (mean 0.48 mm), EW 0.38–0.43 mm (mean 0.40 mm), EI 1.13–1.25; humeral calli distinct, basal impressions short but well developed; apices separately rounded. Punctures inconspicuous; setae much shorter than those on pronotum, sparse, suberect.

Hind wings long, functional.

Legs moderately long and slender, unmodified.

Aedeagus (Figs 36–39) elongate, AeL 0.15 mm; as in H. klapperichianus yaeyamanus .

Female unknown.

Distribution. Amami-Ôshima, the largest of Satsunan Islands, NE Ryukyu Archipelago, Japan ( Fig. 44 ). Etymology. Named after Amami Islands that include Amami-Ôshima.

Remarks. Himaloconnus klapperichianus amamianus has adults distinctly smaller than those of H. klapperichianus klapperichianus and H. klapperichianus yaeyamanus , but comparable in length with those of H. klapperichianus okinawanus. The eyes of males are slightly longer than tempora in dorsal view (they should be measured to make sure), and differ from the eyes of H. klapperichianus okinawanus , which are clearly shorter than tempora. Moreover, the ommatidia of H. klapperichianus amamianus are largest in relation to the head among all subspecies of H. klapperichianus .

Figures

FIGURE 44. Distribution of Himaloconnus in Japan.

FIGURES 1–3. Dorsal habitus of males. Himaloconnus klapperichianus yaeyamanus ssp. n. (1); H. klapperichianus amami- anus ssp. n. (2); and H. klapperichianus okinawanus ssp. n. (3).

FIGURES 4–9. Head of males in dorsal (4, 6, 8) and lateral (5, 7, 9) view. Himaloconnus klapperichianus yaeyamanus ssp. n. (4–5); H. klapperichianus amamianus ssp. n. (6–7); and H. klapperichianus okinawanus ssp. n. (8–9). Vertical lines to the right of each image show length of eye and length of temple.