Bellatrachia condoriana (Crosse & Fischer, 1863)

Pall-Gergely, Barna & Neubert, Eike, 2019, New insights in Trichochloritis Pilsbry, 1891 and its relatives (Gastropoda, Pulmonata, Camaenidae), ZooKeys 865, pp. 137-154 : 140

Bellatrachia condoriana (Crosse & Fischer, 1863) Figs 1-2 , 3-4

Helix condoriana Crosse & Fischer, 1863: 351, pl. 14, fig. 1.

Chloritis microtricha Möllendorff, 1898: 71. syn. nov.

Chloritis (Trichochloritis) microtricha : Zilch 1966: 304, pl. 9, fig. 23.

Trichochloritis microtricha : Schileyko 2011: 47.

Trichochloritis condoriana : Schileyko 2011: 47.

Bellatrachia pseudomiara : Schileyko 2018: 169-171, figs 1-2 [non Helix (Chloritis) pseudomiara Bavay & Dautzenberg, 1909].

Type specimens.

condoriana : 1 syntype MNHN-IM-2000-1866, Poulo-Condor, D: 18.3 mm, H: 11.7 mm [this is the syntype figured in the original description]; microtricha : lectotype (selected by Zilch 1966) SMF 8540, Vietnam, Annam, Boloven, coll. O. Möllendorff ex coll. Roebelen, D: 24.8 mm, H: 17.7 mm.

Type locality.

"insula Poulo-Condor" [Con Son Island], Vietnam.

Additional specimens.

Vietnam, Bang-Kiang, coll. Denis ex Messager, MNHN-IM-2012-27151 (2 shells).

Diagnosis.

Shell biconvex with a whitish subsutural spiral, narrow umbilicus, and hair scars covering the entire surface.

Description.

Shell middle sized, biconvex, moderately thin-walled; last whorl only slightly expanding and descending abruptly towards aperture; colour dirty yellowish with a broad pale subsutural spiral band; whorls 4.5-5, separated by a rather shallow suture; body whorl faintly slightly angled; subsutural furrow shallow but present on the complete last whorl; protoconch consists of 1.25-1.5 whorls, very finely squamous, matte; the pattern of hair scars is dense and covers the complete teleoconch; aperture obliquely rounded, and the peristomal rims are close; peristome strongly expanded and somewhat reflected and reinforced by a white lip; parietal side with very thin, inconspicuous light layer; umbilicus open, of medium size, with blunt peripheral angulation, and partly covered by the columellar reflection.

Measurements.

D = 18.3-24.8 mm; H= 11.7-17.7 mm (n = 4).

Remarks.

The syntype of B. condoriana ( Fig. 1 ) is similar to the specimen identified as Helix (Chloritis) pseudomiara by Schileyko (2017) ( Fig. 3 ), but the shell of the latter is somewhat more depressed. The shell of the lectotype of B. microtricha ( Fig. 2 ) is larger and somewhat more globular that that of B. condoriana . However, both taxa agree quite well in other details such as the relative size of the umbilicus, formation of lip and aperture, and microsculpture of the teleoconch. In contrast, absolute dimensions proved to be insufficient traits for species-level distinction. Therefore, we consider Chloritis microtricha as a synonym of Bellatrachia condoriana . The subtle conchological differences in the shell morphology shown in Figs 1 - 3 may be part of the overall variation of B. condoriana or may signal a difference at the species level. This question can only be clarified by a revision of a larger number of specimens from the area.

Figures

Figures 1 - 2. Bellatrachia condoriana 1 Syntype Helix condoriana Crosse & Fischer, 1863, MNHN-IM- 2000 - 1866, D = 18.3 mm, MNHN 2 lectotype Chloritis microtricha Moellendorff, 1898, SMF 8540, D = 24.8 mm, S. Hof, Senckenberg. All photographs x 2.

Figures 3 - 4. Original specimen " Bellatrachia pseudomiara " sensu Schileyko 3 shell (Schileyko 2018: 170, Fig. 1), D = 23.2 mm, A. Sysoev, photograph x 2 4 Morphology of the genital organs of " Bellatrachia pseudomiara " sensu Schileyko (2018), modified after Schileyko (2018).