Opsiphanes mutatus Stichel, 1902

Piovesan, Mônica, Casagrande, Mirna Martins & Mielke, Olaf Hermann Hendrik, 2022, Systematics of Opsiphanes Doubleday, [1849] (Lepidoptera: Nymphalidae, Satyrinae, Brassolini): an integrative approach, Zootaxa 5216 (1), pp. 1-278 : 111-112

Opsiphanes mutatus Stichel, 1902

( Figs 92–94 )

Opsiphanes sallei mutatus Stichel, 1902 . Berl. ent. Ztschr. 46 (4): 514, 524, fig. 2 (male d); 4 males, XI–XII, S[an]ta Inéz, [Tungurahua], Ecuador eastern Andes, 1,250 m, R. Haensch leg.— Stichel, 1904, in Wytsman. Gen. Ins. 20, p. 20.— Röber, 1906. Soc. ent. 21 (4): 27.— Stichel, 1907. Soc. ent. 22 (13): 100.— Stichel, 1909. Das Tierreich 25: 81, fig. 20 (male d).— Fruhstorfer, 1912, in Seitz. Gross-Schmett. Erde 5, p. 301.— Stichel, 1932. Lep. Cat. 51, p. 43.— Biezanko & Ruffinelli, 1962. Rev. Fac. Agron., Montevideo, 50: 146.— Blandin & Descimon, 1975. Ann. Soc. ent. France 11 (1): 9.— D’Abrera, 1987. Butt. Neotrop. Reg. 3, p. 410, 411 (figs male d, v).— Blandin et al., 2015. Antenor 2 (1): 115.

Opsiphanes sallei mutato [sic]; Stichel, 1904. Ins.-Börse 21 (1): 6.

Opsiphanes mutatus mutatus ; Bristow, 1991. Zool. Jour. Linn. Soc. 101 (3): 281, 286, fig. 31b (male d, v); LECTOTY- PUS.—Moreno et al., 1998. Marip. Ecuador, p. 155.— Casagrande, 2004. Brassolini , in Lamas (ed.) Checklist: Part 4A, Hesperioidea-Papilionoidea, in Heppner (ed.). Atlas Neotrop. Lep. 5A, p. 204.

Opsiphanes mutatus ; Austin et al., 2007. Bull. Allyn Mus. 150: 15, 17.

Taxonomic history. Considered a subspecies of Opsiphanes sallei , until Bristow (1991), based on the wing pattern, raised it to species rank. This decision was followed by Casagrande (2004). Blandin et al. (2015) noted that Opsiphanes mutatus is exclusively Andean.

Type material. Opsiphanes sallei mutatus Stichel, 1902 was described based on four males, from [Hacienda] S[an]ta Inéz [=Inés], [Tungurahua], Ecuador, eastern Andes, 1,250 m, collected between XI–XII by R. Haensch. Bristow (1991) designated a male as the lectotype, deposited at the NHMUK, with the following labels: / Typus/ Type/ LECTOTYPE / Ecuador Santa Inez e. c. H. Stichel/ O. sallei mutatus Stich. Type/ Rothschild Bequest B. M. 1939-1./. Stichel (1904) referred to figure 2 of the original description of Opsiphanes mutatus as “fig. typ.” Bristow (1991) selected as a lectotype a different specimen from the one illustrated by Stichel (1902, 1909) ( Fig. 94 ). Thus, “fig. typ.” of Stichel (1904) should be interpreted as the “typical” phenotype.

Diagnosis. Opsiphanes mutatus is phenotypically similar to Opsiphanes farrago stat. nov., differing in having the DFW with the oblique band usually twice the width of the DHW submarginal band, and the DHW with the submarginal band usually evident only up to M 3 ( Fig. 92a ). Despite the small number of specimens of Opsiphanes mutatus analyzed (n=3), these characters were constant. Female: Unknown.

Male genitalia. Not examined. Female genitalia. Not examined. Variation. FW, length: male: 45–46 mm (n=2). Natural history. Unknown. Geographic distribution. Ecuador (“Oriente”, [Tungurahua]); Peru ( Río Huallaga ) ( Fig. 93 ). Temporal distribution. Unknown. Etymology. Unknown. Specimens examined. Photos of the lectotypus of Opsiphanes mutatus Stichel, 1902 , and the following specimens: ECUADOR: Ecuador Oriente —no locality, 1 male, ex-coll. F. Johnson, AMNH_IZC 00352840 (AMNH). PERU: no department — Río Huallaga , 1 male, ex-coll. F. Johnson, AMNH_IZC 00352845 (AMNH).

Figures

FIGURE 92. Opsiphanes mutatus Stichel, 1902: Male, dorsal and ventral views: a. Ecuador: (AMNH_IZC 00352840-AMNH) Ecuador Oriente; b. Peru: (AMNH_IZC 00352845-AMNH) Rio Huallaga. White arrows indicate diagnostic characters of the species.

FIGURE 93. Geographical distribution of Opsiphanes mutatus Stichel, 1902.

FIGURE 94. Opsiphanes mutatus Stichel, 1902. Lectotype of Opsiphanes mutatus Stichel, 1902; a. dorsal view, b. ventral view.