Phanerotoma artocornuta, Achterberg, 2021

Achterberg, Cornelis van, 2021, The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen, ZooKeys 1014, pp. 1-118 : 1

Phanerotoma artocornuta sp. nov. Figs 13-15 , 16-27

Type material.

Holotype, ♀ (RMNH), " Yemen: Al Kowd (4054), viii.1999, light trap, A. v. Harten & S. Al Haruri, RMNH’00”. Paratypes: 2♀: Idem, i.-iii.2003; 5♀: Idem, iv.2001; 2♀: Idem, vii.-ix.2001; 2♀: Idem, ix.2003; 1♀: Idem, 17-21.vii.2001; 1♀: Idem, 21-25.viii.2001; 1♀: Idem, 8-12.vii.2001; 1♀: Idem, vi.2002; 1♀: Idem, v.-vi.2000; 1♂: Idem, vii.1999; 1♀, "Yemen: Ar Rujum (5700), 9.iv.-5.vi.2001, Mal. trap, A. v. Harten, RMNH’02”; 1♀, "Yemen: Al Kadan (7501), i.2003, light trap; A. v. Harten & T. Abdul-Haq, RMNH’03”; 4♀: Idem, v.2002; 1♂: " United Arab Emirates, Fujairah (1224), light tr[ap], 5-24.iii.2005, 25°08'N, 56°21'E, A. v. Harten, RMNH’06”.

Diagnosis.

Subapical antennal segments of ♀ rather slender, sixth segment from apex narrowed basally and subapically widened and with small and round protuberances near apex, resulting in a somewhat serrate margin of antenna (Fig. 27 ); antenna of ♀ approx. as long as body and eight-eleventh segments from apex elongate; clypeus comparatively transverse and hardly protruding medio-ventrally (Fig. 24 ); parastigma yellow and comparatively large (Fig. 16 ); scutellar sulcus narrow to rather wide; parastigma, pterostigma and all veins pale yellow; antenna of ♀ 1.0-1.1 × as long as body; second tooth of mandible 0.4-0.5 × as long as apical tooth; maximum width of clypeus 0.8 × minimum width of face; temple densely striate and rather shiny; clypeus with 3 minute teeth; length of malar space 0.7-0.8 × basal width of mandible; vein cu-a of fore wing distinctly inclivous; vein r 0.9-1.7 × vein 3-SR and forming an angle (Fig. 16 ); third metasomal tergite 1.4-1.5 × as long as second tergite and with curved sides; hypopygium without up curved triangle or spine apically (Fig. 19 ); length of fore wing 1.8-2.4 mm. Phanerotoma artocornuta is similar to P. longiradialis because of similar antenna, inner tooth of mandible and not angled veins r and 3-SR. P. artocornuta differs by having apical third of antenna pseudo-serrate (cylindrical in P. longiradialis ), anterior tentorial pits distinctly above lower level of eyes (at lower level of eyes), temples directly narrowed behind eyes (roundly narrowed), pterostigma yellow (dark brown) and vein r approx. as long as vein 3-SR (distinctly shorter).

Description.

Female, holotype, length of body (excluding ovipositor) 3.1 mm; antenna 2.5 mm; fore wing 2.3 mm; visible part of ovipositor sheath 0.35 mm.

Head. Width 1.6 × median length in anterior view and part of head above eye in lateral view 0.35 × height of eye (Fig. 25 ); antenna with 23 segments and 1.1 × as long as fore wing, segments near apical quarter of antenna elongate and longer than wide, widened subapically because of small round protuberances and seven apical segments rather moniliform (Fig. 27 ) and apical segment with distinct spine; third, fourth, and penultimate segments 3.8, 3.4 and 2.0 × longer than wide in lateral view, respectively; area of stemmaticum coriaceous; OOL: diameter of posterior ocellus: POL = 12: 4: 5; length of eye 4.0 × temple in dorsal view (Fig. 23 ); frons rugose laterally and densely rugulose medially, and rather shiny, with median carina posteriorly; vertex coarsely transversely rugose-striate and with satin sheen; temple largely striate and rather shiny, its median width 0.6 × width of eye in lateral view; face finely reticulate-rugose and with median ridge dorsally, but no distinct median carina; clypeus mostly smooth, shiny, 0.8 × wider than minimum width of face (intertentorial width 2.2 × minimum distance between clypeus and eye) and with three minute teeth medio-ventrally (Fig. 24 ); eye large, strongly convex and in lateral view 1.3 × (measured medially) as wide as temple (Fig. 25 ), in anterior view its height equal to minimum width of face; upper condyle of mandible below lower level of eyes (Fig. 24 ); malar space rugulose, with satin sheen and 0.7 × basal width of mandible; lower tooth of mandible 0.5 × as long as apical tooth (Fig. 22 ).

Mesosoma (Figs 13 , 17 ). Length 1.8 × its width in lateral view; side of pronotum only medially and posteriorly rugose, remainder rugulose or superficially coriaceous; propleuron posteriorly weakly convex; mesosternum densely granulate and rather matt; mesoscutum densely rugulose; scutellum flat, densely granulate but smooth posteriorly and with satin sheen; notauli not indicated; scutellar sulcus wide and with eight carinae (Fig. 17 ); metanotum without short median carina anteriorly and some micro-sculpture posteriorly; propodeum coarsely rugose-reticulate, without distinct median and transverse carinae, and latero-posteriorly slightly tuberculate. Wings. Fore wing 3.0 × longer than its maximum width; 1-R1 as long as pterostigma; distance between wing apex and 1-R1 0.3 × length of vein 1-R1; r issued distinctly beyond middle of pterostigma, angled to 3-SR and 1.6 × 3-SR; 2-SR weakly curved and distally converging to posterior margin of pterostigma (Fig. 16 ); SR1 straight; 2-SR+M short, m-cu slightly postfurcal; parastigma rather large; first discal cell of fore wing much higher than first subdiscal cell; 1-CU1 0.45 × as long as vein 2-CU1, cu-a 0.9 × 1-CU1; r:3-SR:SR1 = 8:5:47; 2-SR:3-SR:r-m = 19:5:7; r-m nearly vertical; 2-M slightly curved (Fig. 16 ). Hind wing: M+CU:1-M:1r-m = 26:21:10. Legs. Hind femur with satin sheen, 3.3 × as long as wide and rather widened submedially; hind tibia rather swollen; middle tibia with medium-sized yellowish blister; inner spur of middle tibia 0.5 × its basitarsus; hind coxa superficially granulated and shiny.

Metasoma (Figs 18 , 19 ). Elliptical in dorsal view, 1.9 × as long as wide and 1.1 × as long as mesosoma; first and second tergites densely and rather coarsely longitudinally rugose; metasomal sutures medium-sized; third tergite convex medially, 1.4 × longer than second tergite and laterally curved, in lateral view rather convex, largely densely reticulate-rugulose and with satin sheen (Fig. 19 ), lateral lamella narrow laterally, posteriorly rather wide and not protruding latero-apically, medio-apically truncate; ovipositor sheath widened apically, its visible part 0.1 × as long as fore wing and 0.3 × metasomal carapace and its setose apical part with medium-sized setae and 0.05 × as long as fore wing; hypopygium setose and acute apically, without up curved triangle apically or apical spine (Fig. 19 ).

Colour. Pale yellowish; apical antennal segments apically and apex of ovipositor sheath brown; stemmaticum blackish; telotarsi brownish yellow; veins (including 1-M), parastigma and pterostigma pale yellowish; wing membrane subhyaline.

Male.

Very similar to female, but subapical antennal segments more elongate and with an erect apical bristle (Fig. 14 ), hind femur somewhat widened (Fig. 15 ), vein 1-M, parastigma and pterostigma partly dark brown or brown.

Variations.

Length of fore wing 1.8-2.4 mm; inner tooth of mandible rather robust and 0.4-0.5 × as long as apical tooth; vein r of fore wing 0.5-1.7 × vein 3-SR.

Biology.

Unknown.

Distribution.

United Arab Emirates, Yemen.

Etymology.

From artus (Latin for narrow) and cornutus (Latin for horned) because of the slender antenna.

Figures

Figures 13 - 15. Phanerotoma artocornuta van Achterberg, sp. nov. ♀, holotype (but 14 and 15 of ♂, paratype) 13 habitus lateral 14 apical third of antenna lateral 15 hind femur and tibia lateral.

Figures 16 - 27. Phanerotoma artocornuta van Achterberg, sp. nov., ♀, holotype (but 21 of ♀ paratype) 16 wings 17 mesosoma dorsal 18 first-third metasomal tergites dorsal 19 metasoma lateral 20 hind leg lateral 21, 22 mandible ventral 23 head dorsal 24 head anterior 25 head lateral 26 antenna lateral 27 apical half of antenna lateral.