Sulciclivina andrewesi, Balkenohl, 2023

Balkenohl, Michael, 2023, Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini), European Journal of Taxonomy 915 (1), pp. 1-57 : 46-48

Sulciclivina andrewesi sp. nov.

urn:lsid:zoobank.org:act:DF22D598-5910-4EA2-B4F1-6A5EF2506E7B

Figs 23 , 52–53

Diagnosis

The species is the distinctly largest-sized one of the sagittaria- species group. Unique characters are in addition the elongated and falcate mandibles, the elongated and parallel-sided pronotum, and the elongate elytron with the lateral margins parallel.

Etymology

The name is a patronym in honour of Herbert Edward Andrewes (1863–1950) who provided a solid basis for the investigation of Asian Scaritinae .

Type material

Holotype

INDIA • ♀; with labels and data: white, black printed “ 8. IX. 17 1. Among Dabui Rarhi and Batri roots Pusa, Haq. Coll:” / “Agric. Res. Inst. Pusa” / handwritten in black ink and printed “ Clivina sagittaria Bates (over) H.E.Andrewes det.” / “ H.E.Andrewes Coll. B.M.1945-97.”; NHMUK.

Remark. In the left hind leg, all tarsomeres are missing and the right elytron shows a pinhole and a crack. The mandibles are not as sharp as in the paratype due to individual wear.

Paratype

INDIA • 1 ♀; with labels and data: white, black printed and handwritten in black ink “2110 Pusa Behar At Light” / “Agric. Res. Inst. Pusa” / “ H.E.Andrewes Coll. B.M.1945-97.”; NHMUK .

Remarks. The left elytron and the left hind leg are missing, the right elytron shows a pinhole. The ventral surface shows cracks and in the left intermediate leg, three tarsomeres are missing. Andrewes (1929: 359) already mentioned some of the outstanding characters of these two specimens but did not describe them as a new taxon.

Description

MEASUREMENTS. Holotype / paratype: body length 6.94/ 6.57 mm, width 1.57 / 1.61 mm; L/W of pronotum 1.2 / 1.81; L/W of elytra 2.31 / 2.23.

COLOUR. Glossy. Piceous. Mouthparts, antenna, elytron, intermediate and hind leg, tarsomeres of front leg fuscous, palpomeres more pale fuscous. Anterior part of supraantennal plate at the margin slightly translucent-fuscous.

HEAD. A quarter narrower than pronotum. Clypeus with central part distinctly bilobed anteriorly; lateral lobe developed as acute tooth, nearly as projecting as central lobes, clypeal wing well developed, rounded, not as far projecting anteriorly as lateral tooth of clypeus, separated from lateral lobe by obtuse notch. Supraantennal plate laterally moderately convex, laterally somewhat less projecting than eye, indistinctly reflexed margined. Supraantennal plate overlapping eye anteriorly by about a fifth (dorsal view), with the supraorbital carina keeled posteriorly and extended up to mid-eye level, separated from wing by sharp notch. Clypeus transverse, separated from frons by sharp carina in form like an obtuse inverted V. Frons distinctly convex, with another but smaller carina like an inverted V which encloses elevated area, supraorbital carina short and tubercle-like, extending up to posterior gena-level. Clypeus rugose, area between V-like carinae with transverse rugae, supraantennal plates and frons rugose, posteriorly with distinct longitudinal rugae, clypeus and frons separated from supraantennal plate and supraorbital carina by deep and moderately wide furrow; with two supraorbital setigerous punctures situated at posterior eye and at end of gena-level. With some moderately sized punctures laterally posterior eye. Eye small, embedded, moderately convex in dorsal view, main part situated ventrally where it is more convex. Gena moderately convex, covering around a sixth of posterior eye in ventral view. Eye somewhat constricted due to the overlapping supraantennal plate and the genae. Antenna not reaching up to middle of pronotum, antennomeres five to ten moniliform (L/W around 1.1). Labrum straight anteriorly, with isodiametric-irregular reticulation, six setose. Mandibles distinctly falcate, bent up dorsally. Mentum with irregular fine wrinkles and indistinct isodiametric reticulation, shape of lobe wide-oval, flattened, with sub-marginal carina, with long median carina. Median tooth moderately wide, not as protruding anteriorly as lobe, hollowed out, obtuse angled and bicarinate anteriorly.

PRONOTUM. Disk slightly convex in lateral and semicircular in frontal view. Distinctly longer than wide, peltate. Reflexed lateral margin nearly smooth, straight laterally, nearly parallel, anterior angle rounded off, posterior angle missing; lateral channel moderately narrow, narrower before and behind posterior setigerous puncture. Median line moderately deep, sharp, subcrenulated, complete. Anterior transverse crenulated, wider than median line, joining median line, joining declivity at anterior margin. Surface glossy, completely covered with small punctures, with paralateral indistinct group of moderately sized punctures, with few fine transverse wrinkles laterally and basally. Base distinct, channel two times as wide as lateral channel, width about a quarter of width of pronotum. Proepisternum distinct, well visible in dorsal view.

ELYTRON. Shape elongate with margin nearly parallel. Disk flattened in anterior half in lateral view, semicircular in frontal view. More than two times longer than wide, with maximum at middle. Reflexed lateral margin smooth, setigerous punctures in lateral channel with regularly long setae, punctures widely interrupted at middle. Scutellar striole missing; setigerous tubercle at base of first stria, with tubercle at base of third interval, basal declivity with isodiametric reticulation. Humeral tooth situated at base of fourth interval. All striae distinctly deepened, punctuate-striate, one to three free at base, four reaching humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals distinctly convex, more convex laterally. Setigerous punctures missing on intervals. Surface of all intervals glossy, interval four to eight with indistinct reticulation at humerus, interval eight at apex sub-carinate and with indistinct reticulation.

HIND WINGS. Alae fully developed (based on two specimens).

LOWER SURFACE. Proepisternum covered in lateral half with dense and deep transverse wrinkles, with some moderately sized punctures, at the lateral margin with isodiametric reticulation. Sternite of abdomen smooth at middle but with micropunctures, covered laterally with moderately large punctures.

LEGS. Protibia covered with longitudinal reticulation, with three spines of moderate length, apical spine distinctly arcuate, movable spur robust, regularly arcuate.

MALE GENITALIA. Unknown.

FEMALE GONOCOXITE AND EPIPLEURITE (Fig. 52). Gonocoxite fir-cone shaped, flattened dorsally and ventrally, with three wide nematiform setae positioned in the apical half at the median side, with one preapical nematiform seta. Epipleurite semi-square, with long widened angulated rod.

Variation

The carina on the frons of the head is arch-like rounded in the HT and obtuse-angled in the PT. The punctures on disk of the pronotum are different in intensity and density between the two specimens.

Sexual dimorphism

Unknown.

Distribution

Known from the type locality Pusa, Bihar, in India.

Figures

Figs 20–23. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 20. S. sagittaria sagittaria (Bates, 1892), ♂, paralectotype (NHMUK). 21. S. sagittaria singaporensis subsp. nov., ♀, holotype (NHMUK). 22. S. oculiangusta sp. nov., ♀, holotype (NHMUK). 23. S. andrewesi sp. nov., ♂, holotype (NHMB).