Sulciclivina karelkulti karelkulti, Balkenohl, 2023

Balkenohl, Michael, 2023, Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini), European Journal of Taxonomy 915 (1), pp. 1-57 : 36-38

Sulciclivina karelkulti karelkulti sp. nov.

urn:lsid:zoobank.org:act:2776C7B3-A68C-42D3-ADAF-DB6FDA5162E8

Figs 18 , 48, 53

Diagnosis

A medium-sized species with a long-oval shape of the elytron, with extensive reticulation on the intervals of the elytron, and fused clypeal wings with the lateral lobes. Distinguished from the other species by the reticulation on the elytron, which covers laterally intervals six to eight, apically the whole apex of the elytron by one fifth, and basally it is extended on intervals one and two. The gonocoxites are pointed apically but not elongated at the apex as it is in the similar species S. curvata sp. nov.

Etymology

The name is a patronym in honour of the Czech entomologist Karel Kult, specialist in Scaritinae , who worked intensively on Clivinini between 1940 and 1960.

Type material

Holotype

MALAYSIA • ♀; with label and data: beige, handwritten in black ink “Malacca Prof. Fischer”; MFNB.

Paratypes

MALAYSIA • 7 ♀♀; “MALAY PENIN. WEST COAST.; LANGKAWI IS.; 14, 15, 16, and 21 Apr. 1928 ” / “H. M. Pendlebury Coll. F.M.S. Museums.” / “Ex F.M.S. Museum. B.M.1955-354.”; NHMUK • 1 ♀; “Pekan PAHANG 10. IX. ’17 G.D.A.” / “Ex Mus. Coleoptera 34 Hope Ent. Coll. Oxford”; OUMNH • 1 ♀; “ MALAYSIA, Pahang 40km W Rompin Selendang, 29.4.- 6.6.1993 leg. I.Jenis ”; NHMW • 1 ♀; “ Kota Tinggi Jahore August 1917 ” / “ Clivina attenuata Herbst H.E.Andrewes det.” / “OX. UNI. MUS. NAT. HIST (OUMNH) G.Dexter-Allen Coll. ”; OUMNH • 1 ♀; “ Jahore. April. H.N.Ridley. 99- 127.” / “ H.E.Andrewes Coll. B.M.1945-97”; NHMUK • 1 ♀; “Prac. Malacca” / “ Clivina attenuata-group det. M. Balkenohl,2017 ”; MFNB • 1 ♀; “ Sungkei 6.2.02 ” / “Ex Misc. Coleoptera 34 Hope Ent. Coll. Oxford ”; OUMNH .

Remark. In the holotype, the right eight terminal antennomeres and the four terminal tarsomeres of the front leg are missing.

Description

MEASUREMENTS. Holotype: body length 6.66 mm, width 1.94 mm; L/W of pronotum 0.99; L/W of elytra 1.86. All material: body length 6.65–7.56 mm (ẍ: = 7.04 mm *), width 1.94–2.14 mm (ẍ: = 2.02 mm *), L/W of pronotum 0.98–1.02 (ẍ: = 1.0*), L/W of elytra 1.83–1.89 (ẍ: = 1.86*); (*n = 7).

COLOUR. Glossy. Piceous. Labrum, intermediate and hind leg fuscous, antenna pale fuscous. Supraantennal plates fuscous, translucent at the margin.

HEAD. A third narrower than pronotum. Clypeus with central part distinctly bilobed anteriorly; lateral lobe prominent, somewhat more protruding anteriorly than central part, with lateral lobe and wing fused, with longitudinal carina at middle. Supraantennal plate longitudinally convex laterally, laterally less projecting than eye. Lateral lobe slightly reflexed margined. Supraantennal plate distinctly reflexed margined. Supraantennal plate overlapping eye anteriorly by a fifth (dorsal view), with supraorbital keel ending before mid-eye level, separated from wing by obtuse-angled notch. Clypeus transverse, distinctly convex, separated from frons by distinct sharp step and wide furrow-like sulcus, step and sulcus in form like an inverted flat V. Frons moderately convex, with distinct arch-like impression at middle, which is step-like impressed laterally, with supraorbital carina up to posterior eye-level. Clypeus smooth, frons with few minute scattered punctures, separated from supraantennal plate and supraorbital carina by deep moderately wide furrow, furrow with small oblique carina posterior mid-eye level; with two supraorbital setigerous punctures, one situated posterior mid-eye level and one at the posterior end of the gena. Neck constriction missing. Eye distinctly convex in dorsal view, projecting laterally; main part of eye situated ventrally where it is globose; eye with narrow but distinct carina situated directly at the median margin. Due to the overlapping supraantennal plate and the gena, the eye looking in lateral view constricted. Gena distinct, indistinctly convex, covering a quarter of posterior eye in ventral view. Antenna relatively short, just reaching over the anterior third of the pronotum, antennomeres five to ten moniliform (L/W around 1.04). Labrum indistinctly trilobed anteriorly, with isodiametric reticulation, six setose. Mentum with isodiametric reticulation, with lobe flattened, shape of lobe asymmetric oval; distinctly carinate at middle, median tooth slightly dilated to apex, not as protruding anteriorly as lobe, slightly hollowed out and obtusely rounded anteriorly.

PRONOTUM. Disk slightly convex in lateral view, in frontal view regularly convex. As long as wide. Reflexed lateral margin smooth, slightly convex with straight part in anterior half, widest behind middle; anterior angle slightly projecting, posterior angle missing; lateral channel narrow, not deeply embedded, indistinctly reticulated, completely and regularly rounded off at level posterior setigerous puncture and continuing narrowly up to base. Median line moderately wide, complete. Anterior transverse line appearing as line, joining median line, just not joining anterior margin. Surface glossy, basolaterally with group of impressed punctures of moderate size, with distinct paramedian group of punctures in basal half, with few transverse wrinkles, disk with few microscopic punctures. Base distinctly marked, three times as wide as lateral channel. Proepisternum indistinct, just visible posteriorly in dorsal view.

ELYTRON. Disk slightly convex in lateral view, distinctly convex in frontal view with increasing convexity towards lateral margin. Distinctly less than twice as long as wide, outline long-oval, with short straight part posterior humerus, with maximum width posterior middle. Reflexed lateral margin smooth. Scutellar striole missing; setigerous tubercle at base of first stria, with small but distinct tubercle at base of second stria. Humeral tooth situated at base of fourth stria. Striae moderately deep, moderately punctuate-striate, one to three free at base, four reaching humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals moderately convex, more convex laterally, interval eight with carina apically. Setigerous punctures missing on intervals. Surface of intervals glossy, interval eight and seven completely, and interval six partly covered with isodiametric reticulation, base with reticulation, extending caudally on interval one and two. All intervals covered with reticulation at apical fifth starting on interval one in apical third (see also under variation).

HIND WINGS. In the 13 female specimens available the alae are fully developed.

LOWER SURFACE. Proepisternum covered in lateral two thirds with isodiametric reticulation and with distinct transverse wrinkles, medially in addition with punctures. Sternite of abdomen fully reticulated from the basal to the apical margin, laterally with moderately large and dense punctures, apical segment completely covered with moderately large punctures; in females in addition with isodiametric reticulation laterally at the third and fourth sternite.

LEGS. Tibiae covered with isodiametric reticulation. Protibia with three spines of moderate length, not sulcate dorsally, movable spur slightly arcuate apically.

MALE GENITALIA. Unknown.

FEMALE GONOCOXITE AND EPIPLEURITE (Fig. 48). Gonocoxite less sclerotized, monomeric, elongate foliform, flattened dorsally and ventrally, with four long nematiform setae positioned at the median side, with one very long nematiform seta situated at the pointed apical tip. Epipleurite oblong-rectangular, with moderately widened angulated rod.

Variation

In all specimens, intervals six to eight of the elytron are covered with distinct isodiametric reticulation. From the base, the apex, and the lateral intervals, the reticulation extends to the disk more or less intensively. It becomes less distinct towards the disk. In one of the specimens, the disk is covered with indistinct reticulation. Some of the specimens show minute scattered punctures on the frons of the head.

Sexual dimorphism

Unknown.

Distribution

Known to occur on the Malayan peninsula.

Figures

Figs 12–19. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 12–13. S. striata striata (Putzeys, 1846). 12. ♂, lectotype, [specimen not cleaned], (OUMNH). 13. ♂ from Bandra, India (NHMUK). 14. S. striata kottea subsp. nov., ♀, holotype (NHMUK). 15. S. curvata sp. nov., ♂ from Java (ETHZ). 16–17. S. sulcigera (Putzeys, 1867). 16. ♂ from Nakhon, Thailand (NHMUK). 17. ♀ from Siem Reap, Cambodia, showing the reduced alae (RBINS). 18. S. karelkulti karelkulti sp. nov., ♀, holotype (MFNB). 19. S. karelkulti medanensis subsp. nov., ♀, holotype (SMNS).