Sulciclivina splendida, Balkenohl, 2023

Balkenohl, Michael, 2023, Revision of the genus Sulciclivina Balkenohl, 2022 from Asia (Coleoptera, Carabidae, Clivinini), European Journal of Taxonomy 915 (1), pp. 1-57 : 23-25

Sulciclivina splendida sp. nov.

urn:lsid:zoobank.org:act:3D27B342-B894-4710-98CF-487D261D1D20

Figs 2 , 11 , 30, 42, 53

Diagnosis

A small-sized species with long-oval elytra with straight and diverging part of the lateral margin, all intervals of elytra smooth, and the lateral lobe of the clypeus completely fused with the wing. Distinguished from the most similar species S. basiangusta sp. nov. by the much less convexity of the elytron, the completely missing clypeal wing, and the regularly sized base of the pronotum. It differs from S. coxisetosa sp. nov. in the long-oval elytron with straight and diverging part of the lateral margin, the different puncture pattern on the disk of the pronotum, and the different shape of the gonocoxites.

Etymology

The name refers to the smooth and glossy interval eight and apex of the elytron by the Latin adjective for smooth and shiny in the feminine form ( ʻ splendida ʼ).

Type material

Holotype

INDIA • ♂; with white black printed and handwritten with black ink label and data: “INDIA Barway” [INDIA Barway = India, Chattisgarh state, Jashpur district , Chainpur env.]/ “ C. andrewesi Kult 47 det. K. Kult ” / red, black framed and black printed “TYPE” / white, black printed and black handwritten in black ink “in litteris V. att.-Gr. det. Dr. A. Dostal 2000” / “COLLECTIO KAREL KULT COLL. A.DOSTAL, 1999” / white, black printed “Coll. Dostal”; CADW.

Remark. The following parts are missing: apical segment of the left antennae, tarsomeres two to five and movable spur of the left front leg, tarsomeres four and five of the left intermediate leg, and complete right hind leg. The specimen was received with empty hind body with the median lobe of the aedeagus dry mounted beside the specimen and without parameres.

Paratypes

INDIA • 1 ♀; with white handwritten with black ink and black printed labels and data: “Nagpore” / “T.R.S.N.COLL.” / “ Coll. J. Saltin in Coll. CBA 1997”; CMBB • 1 spec.; “Pegu India ” / “Bought from Staudinger & Bang – Haas, 1923” / “ Clivina andrewesi Kt. det. K. Kult, 1948” / “Compared with TYPE K. Kult, 1948” / “ H.E.Andrewes Coll. B.M. 1945-97.”; NHMUK .

Remark. The following parts are missing: segments three to eleven of the left antennae. The second specimen was received with an empty abdomen, obviously due to a dissection attempt. The two hind legs, two terminal antennomeres of the right antennae, and the three terminal tarsomeres of the intermediate legs are missing.

Taxonomic remark

Kult already recognized the form as being a new species and added a red type label and a name to the specimens. However, it was never described and the name was never published. The name assigned to the specimen by Kult is therefore not valid for this form.

Description

MEASUREMENTS. Holotype: body length 5.15 mm, width 1.57 mm; L/W of pronotum 0.95; L/W of elytra 1.82. Paratype: body length 5.61 mm, width 1.67 mm; L/W of pronotum 0.94; L/W of elytra 1.84.

COLOUR. Glossy. Piceous. Labrum, intermediate and hind leg fuscous, antenna and tarsomeres of front leg fuscous. Supraantennal plate at the margin and anterior part of lateral margin of pronotum translucent-fuscous.

HEAD. A third narrower than pronotum. Clypeus with central part nearly straight, indistinctly bilobed anteriorly; lateral lobe distinctly projecting, wing completely fused with lateral lobe. Supraantennal plate laterally less convex at middle, laterally somewhat less projecting than eye; clypeus and wing slightly reflexed margined, supraantennal plate distinctly reflexed margined. Supraantennal plate overlapping eye anteriorly by less than a fifth (dorsal view), with supraorbital keel extended nearly continuously up to gena level, separated from wing by sharp notch. Clypeus transverse, slightly convex, separated from frons by distinct sharp sulcus, sulcus in form of convex arch. Frons moderately convex, with two small longitudinal impressions at middle, with supraorbital carina up to posterior eye-level. Clypeus and frons with few scattered micro punctures, separated from supraantennal plate and supraorbital carina by deep moderately wide furrow; with two supraorbital setigerous punctures at middle and posterior end of the eye. Neck constriction missing. Eye convex in dorsal view. Main part of eye situated ventrally where it is globose. Due to the overlapping supraantennal plate and the gena, the eye resembles in lateral view compressed. Gena distinct, moderately convex, covering a quarter of posterior eye in ventral view. Antenna moderately short, reaching basal third of pronotum, antennomeres five to ten moniliform (L/W around 1.1). Labrum nearly straight anteriorly, with isodiametric-irregular reticulation, six setose. Mentum with isodiametric reticulation at middle, with lobe flattened, shape of lobe semi-ovoid and gently bent medially, irregularly to isodiametrically reticulated, with short carina at median tooth. Median tooth moderately wide, not as protruding anteriorly as lobe, carinate at base, bluntly rounded anteriorly.

PRONOTUM. Disk moderately convex in lateral and distinctly convex in frontal view. Wider than long, subsquare. Reflexed lateral margin smooth, attenuating in anterior half with straight part, widest behind middle; anterior angle distinct, slightly projecting, posterior angle missing; lateral channel moderately narrow, completely and regularly rounded off before posterior setigerous puncture and continuing up to base. Median line finely engraved, somewhat punctured, complete. Anterior transverse line consisting of longitudinal connected punctures, joining median line, indistinctly joining anterior margin. Surface with micro punctures, with circular group of medium-sized punctures basolateral, with paramedian impression basally, with few transverse wrinkles. Base distinctly marked, three times as wide as lateral channel, with width a quarter of pronotum. Proepisternum well visible in dorsal view but not enlarged.

ELYTRON. Disk slightly convex in lateral view in anterior half, distinctly and regularly convex in frontal view. About 1.8 times as long as wide, shape long-oval with straight and diverging part in anterior half, with maximum width slightly behind middle. Reflexed lateral margin smooth. Scutellar striole missing; setigerous tubercle at base of first stria, with small but distinct tubercle at base of third interval, basal declivity with irregular reticulation. Humeral tooth situated at base of fourth interval. Striae moderately deep, distinctly and regularly punctuate-striate, one to three free at base, four reaching humerus, one and two ending free at apex, three and four, and five and six joining apically. Intervals convex, more convex laterally, interval eight narrow and carinate at apex. Setigerous punctures missing on intervals. Surface of intervals glossy, interval eight smooth, intervals smooth at apex.

HIND WINGS. In the HT, the alae are reduced to a small rudiment, the PT is fully winged.

LOWER SURFACE. Proepisternum laterally with small stripe of indistinct irregular reticulation, with some transverse wrinkles, and with scattered small punctures. Sternite of abdomen laterally with limited number of moderately large punctures and with small wrinkles, smooth posteriorly, smooth at middle but with microscopic punctures, apical segment with medium-sized punctures, small wrinkles, and flat impression laterally in the HT; in the PT with distinctly less punctures.

LEGS. Protibia covered with distinct longitudinal reticulation, with three spines of moderate length, not sulcate dorsally, movable spur apically slightly arcuate. Intermediate tibia nearly smooth.

MALE GENITALIA (Fig. 30). Median lobe robust, arcuate in basal third, in apical two-thirds indistinctly bisinuately elongated; apically with indistinct dorsal bulge, with slightly distorted spoon-like spatula, with no visible microtrichia on the surface. Endophallus longitudinally folded, with group of strong bristles at middle.

FEMALE GONOCOXITE (Fig. 42). Gonocoxite foliform, slender, flattened dorsally and ventrally, with three long nematiform setae positioned at the median side, with one apical nematiform seta arising from the pointed and somewhat distorted apex of the gonocoxite.

Variation

Beside the different development of the alae, the two longitudinal impressions at the middle of the frons are more or less deeply impressed. The HT is somewhat paler in colour and is considered as a slightly immature stage.

Sexual dimorphism

In the female (PT), each of the sternites of the abdomen show apically less punctures than in the male (HT). The apical segment shows in total less punctures in the female.

Distribution

Known from the type localities in Barway and Nagpur, India.

Figures

Figs 1–3.Sulciclivina Balkenohl, 2022, head in dorsal view, showing the specific terms used. 1. Sulciclivina attenuata attenuata (Herbst, 1806), ♀, lectotype (MFNB). 2. S. splendida sp. nov., ♂, holotype (NHMB). 3. S. sagittaria sagittaria (Bates, 1892), ♂, paralectotype (NHMUK).

Figs 4–11. Sulciclivina Balkenohl, 2022, habitus, dorsal view. 4–5. S. attenuata attenuata (Herbst, 1806). 4. ♀, lectotype (MFNB). 5. ♂ from Swat (CMBB). 6. S. attenuata semireticulata subsp. nov., ♂, holotype (MNHN). 7. S. bhamoensis (Bates, 1892), ♀, paralectotype (RBINS). 8. S. mikirensis sp. nov., ♀, holotype (NHMB). 9. S. basiangusta sp. nov., ♂, holotype (NHMB). 10. S. coxisetosa sp. nov., ♀, holotype (NHMB). 11. S. splendida sp. nov., ♂, holotype (CADW).