Paramanota paxillosa, Hippa, Heikki, 2010

Hippa, Heikki, 2010, Review of the genus Paramanota Tuomikoski (Diptera, Mycetophilidae), with the description of new fossil and recent species, Zootaxa 2618, pp. 47-60 : 57-59

Paramanota paxillosa sp. n.

Figs 6 A–D

Male. Colour. Head yellowish brown, vertex and antennal flagellomeres 2–14 darker brown; setae and other vestiture dark. Thorax yellowish brown, scutum and scutellum brown, antero-dorsal part of anepisternum infuscated; thoracic setae mostly worn out, the few existing ones dark. Legs yellowish brown, the setae and other vestiture dark. Wing brownish; haltere yellowish with dark brown knob. Abdomen brown, hypopygium yellowish brown on basal half, setae dark. Head. Antennal flagellomeres 3–5, Fig. 6 A. Maxillary palp similar to fig. 7 a in Hippa et al. (2004), ultimate palpomere 1.8 times longer than the penultimate one. The strong postocular setae are broken in the specimen and could no be counted. Thorax similar to fig. 8 a in Hippa et al. (2004). Legs. Front tibia lost on both sides. Wing similar to Fig. 1 D. Wing length 2.2 mm. Hypopygium, Figs 6 B, C, D: Gonocoxae ventrally separated by a membranous area, each side with a very large lobe which is posteriorly extending further than the gonostylus, with the posterior part shallowly divided into a broader mesial and narrower lateral sub-lobe; the ventral surface evenly covered with rather long setose, the dorsal surface of the broader mesial lobe with an area of numerous megasetae at the mesial margin; the dorsal surface of the narrower lateral lobe with a rounded ridge with numerous megasetae. The part visible in dorsal view of the gonocoxa is simple with setae similar to those of the ventral side. Tergite 9 simple, the posterior margin concave, the posterolateral corner only slightly lobe-like prolonged, setae similar to gonocoxa. Cercus simple. The hypoproct is slightly obscured in the slide and thus is not drawn in Fig. 6 B. There is a setose sclerite latero-ventrad from the hypoproct which cannot be easily observed in the slide and again is not drawn in Fig. 6 B. Gonostylus with a dorsal lobe, a ventral lobe and a curved median lobe; the dorsal lobe in dorsal view elongate subtriangular, tapering towards the apex, with a comb-like row of submembranous pale lamellae along the postero-lateral margin; expanded ventral lobe at apex, with a mesial comb-like row of dark sclerotized lamellae, and on the ventral side of the comb with a plate-like small lobe with four setae at margin; details of the long median lobe not observable; between the base of the latter and the ventral lobe there seems to be an aggregation of dark lamellae on a small lobe but the character is very difficult to see on the slide. Parameres and aedeagus fused, aedeagal apodemes long, directed obliquely anteriod, ejaculatory apodeme not observable in the slide.

Female. Unknown.

Discussion. P. paxillosa is similar to P. peninsulae and P. schachti . It is distinguished from both by having the dorsal lobe of the gonostylus in dorsal aspect widening from base to apex, not the opposite, by having the median lobe of gonostylus as long as the dorsal and the ventral lobes, not only half of their length, and by having the ventral lobe of the gonostylus only slightly longer than broad instead of being several times longer. Furthermore, P. paxillosa differs from P. peninsulae by having the lateral sub-lobe on the ventral gonocoxal lobe shorter than broad, subtriangular, instead of being about twice longer than broad, thumb-like. In this respect P. paxillosa resembles P. schachti , but the mesial sub-lobe is much more weakly pronounced than in the latter. See also under P. s c h a c h t i.

Etymology. The name is Latin, paxillosa , full of pegs, referring to the numerous peg-like megasetae dorsally on the ventral gonocoxal lobes.

Types. Holotype. Male. THAILAND, Nakhon Si Tammarat, Namtok Yong NP, TV aerial, 8o14.262’N 99o48.289’N, 966m, Malaise trap 11–18.viii.2008, Paiboon leg, T3109 (in QSBG).

Figures

FIGURE 1. Wing, dorsal view (A and C – F) and ventral view (B). A, B. Paramanota grandaeva sp n. (holotype). C. P. orientalis Tuomikoski (Thailand). D, F. Paramanota furcillata sp. n. (D holotype, F paratype). E. P. bifalx sp. n. (holotype). Scale 1.0 mm.

FIGURE 6. Paramanota paxillosa sp. n. (holotype). A. Antennal flagellomeres 3 – 5, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. D. Aedeagus with associated structures, ventral view. Scale 0.10 mm.