Quercus virginiana Mill. (Fagaceae)

WEIRAUCH, CHRISTIAN E., 2006, New Genera, New Species, and New Combinations in Western Nearctic Phylini (Heteroptera: Miridae: Phylinae), American Museum Novitates 3521 (1), pp. 1-44 : 7-9

HOST: Breeds on Quercus virginiana Mill. (Fagaceae) , but also on other species of Quercus . Wheeler (1991) recorded H. guttulosus on Quercus illicifolia Wangenh.

DISTRIBUTION (fig. 10): Specimens examined here range in the United States from Arizona in the west to Florida and South Carolina, in the east. The few Mexican records show that the distribution of H. guttulosus extends as far south as the San Luis Potosí state. In addition to records from specimens studied during this project, which are represented in the distribution map, Uhler (1894) recorded H. guttulosus from Baja California, Mexico, Knight (1941) from Illinois, and Froeschner (1949) from Missouri.

DISCUSSION: The holotype of H. guttulosus ( Reuter, 1876) is in the Zoological Museum of the University, Helsinki, and was examined during this project.

Hamatophylus guttulosus is not restricted to Quercus virginiana . In Mexico, it is also known from Q. polymorpha and Q. mircrolepis , among others, and Wheeler (1991) found it on Q. illicifolia in Pennsylvania. Nevertheless, the rather close matching of the distribution of Q. virginiana with the distribution here displayed for H. guttulosus might be an indication that Q. virginiana is an important if not the primary host for this bug species.

MATERIAL EXAMINED: MEXICO: Nuevo Leon: 30 km W of Linares on Rt 58 toward San Roberto, 24.84331 ° N 99.84493 ° W, 23 Mar 1985, R. T. Schuh and B. M. Massie, Quercus microlepis (Fagaceae) , 13, 1♀ Quercus microlepis (Fagaceae) , det. K. Nixon 1985, 13, 1♀ ( AM). Quercus microlepis (Fagaceae) , 103, 7♀ Quercus microlepis (Fagaceae) , det. K. Nixon 1985, 103, 7♀ Quercus virginiana (Fagaceae) , 1♀ Quercus rysophylla Woatherby (Fagaceae) , det. K. Nixon 1985, 23 ( AMNH). Quercus microlepis (Fagaceae) , 13, 1♀ Quercus microlepis (Fagaceae) , det. K. Nixon 1985, 13, 1♀ ( ZISP). 7.5 miles S of Monterrey, 25.55771 ° N 100.31666 ° W, 12 Mar 1976, Vincent and Schaffner, 103, 8♀ ( TAMU); 17 Mar 1975, Clark and Schaffner, 33, 8♀ Quercus sp. (Fagaceae) , 2♀ ( TAMU); 21 Mar 1974, J. C. Schaffner, Quercus polymorpha (Fagaceae) , 83, 4♀ ( TAMU); 26 Mar 1979, J. C. Schaffner, Quercus sp. (Fagaceae) , 43, 1♀ Quercus polymorpha (Fagaceae) , 3♀ ( TAMU). Valle Alto just S of Monterrey, 25.57361 ° N 100.25694 ° W, 17 Apr 1978, Henry, Schaffner, and Schuh, Quercus sp. (Fagaceae) , 1♀ ( AMNH). San Luis Potosı´: 2 miles E of Ciudad del Maiz, 22.39999 ° N 99.56856 ° W, 12 Apr 1979, T. P. Friedlander and J. C. Schaffner, Quercus sideroxyla Humb. & Bonpl. (Fagaceae) , det. K. Nixon 1988, 33, 5♀ ( TAMU). USA: Arizona: Gila Co. : 8 mi SW jct Rts 87 and 188 (off Rt 87), Tonto National Forest, 33.55989 ° N 111.21341 ° W, 1219 m, 27 May 1983, R. T. Schuh and G. M. Stonedahl, Light Trap, 23 ( AMNH). Mohave Co.: Hualapai Mountains, SE of Kingman, T20N R15W, 35.18944 ° N 114.05222 ° W, 1585 m, 9 Jun 1983, R. T. Schuh, M. D. Schwartz, G. M. Stonedahl, Quercus sp. (Fagaceae) , 2♀ ( AMNH). Yavapai Co.: US Rt 17, 2 mi S of Rt 179, 34.53604 ° N 111.93105 ° W, 1219 m, 18 Apr 1982, M. D. Schwartz, Juniperus monosperma (Cupressaceae) , 1♀ ( AMNH). Florida: Alachua Co. : Gainesville, 29.65139 ° N 82.325 ° W, 22 Apr 1952, J. R. Vockeroth, 13 ( CNC); 1 Apr 1960, H. F. Howden, 1♀ ( CNC); 23 Apr 1952, J. R. Vockeroth, 1♀ ( CNC). Gainesville, 29.65139 ° N 82.325 ° W, 12 Mar 1975, H. Greenbaum, Light Trap, 1♀ ( TAMU). Clay Co.: Bellair, 30.17417 ° N 81.74083 ° W, [date and collector unknown], 1♀ ( CAS). 2♀ ( TAMU). Duval Co.: Jacksonville, 30.33194 ° N 81.65583 ° W, [date and collector unknown], 13 ( TAMU). Gulf Co.: Saint Joseph Penninsula State Park, 28.37556 ° N 82.28694 ° W, 23 Apr 1998, P. Kovarik, Quercus sp. (Fagaceae) , 13, 1♀ ( TAMU). Highlands Co.: Archbold Biological Station, 27.18833 ° N 81.33778 ° W, 23 Apr 1967, D. E. Bright, 13 ( CNC). Hillsborough Co.: Tampa, 27.94722 ° N 82.45861 ° W, 24 Mar 1986 – 26 Mar 1986, J. E. Eger, 173, 7♀ ( TAMU); 20 Apr 1986, J. E. Eger, 1♀ ( TAMU); 3 Apr 1986, J. E. Eger, 13 ( TAMU). Lake Co.: Hawthorne, 28.76111 ° N 81.87111 ° W, 2 Jun 1961, L. A. Kelton, 113, 7♀ ( CNC). Levy Co.: 3.8 mi S of Bronson P.O., 29.39232 ° N 82.6425 ° W, 29 Apr 1984, J. T. Polhemus, 13, 2♀ ( JTP). 4.5 mi NE of Cedar Key on rt. 24, 29.18452 ° N 82.98238 ° W, 29 Apr 1984, J. T. Polhemus, 2♀ ( JTP). Liberty Co.: TNC Apalachicola Bluffs & Ravines Pres., Travelers Tract, 29.72556 ° N 84.98333 ° W, 14 May 1999, P. W. Kovarik, Quercus sp. (Fagaceae) , 13 ( TAMU). Torreya State Park, 30.56889 ° N 84.94778 ° W, 19 Apr 1984, C. B. Barr, Light Trap, 13 ( UCB). Manatee Co.: Oneco, 27.44722 ° N 82.54639 ° W, 22 Mar 1955, J. C. Martin, 13 ( CNC). Miami-Dade Co.: Florida City, 25.4475 ° N 80.47944 ° W, 28 Mar 1939, G. B. Merrill, Light Trap, 2♀ ( USNM). Kendall, 25.67889 ° N 80.3175 ° W, 28 Apr 1961, L. A. Kelton, 1♀ ( CNC). Orange Co.: Winter Park, 28.59972 ° N 81.33944 ° W, 26 Mar 1111, E. M. Davis, 1♀, Light Trap, 13 ( USNM); 11 Apr 1111, E. M. Davis, 13, 1♀ ( USNM); 28 Mar 1111, E. M. Davis, 13 ( USNM); 25 Mar 1111, E. M. Davis, 1♀ ( USNM); 15 Mar 1944, H. T. Fernand, Light Trap, 1♀ ( USNM). Volusia Co. : Port Orange, 29.13806 ° N 80.99583 ° W, 23 Mar 1963 – 8 Apr 1963, J. F. Brimley, 33, 1♀ ( TAMU). South Daytona, 29.16556 ° N 81.00472 ° W, 3 Apr 1961, J. F. B., 1♀ ( TAMU). Georgia: Glynn Co.: Saint Simons Island, 31.15308 ° N 81.37464 ° W, 22 Apr 1911 – 12 May 1911, J. C. Bradley, 1♀ ( CAS). Liberty Co.: Saint Catherines Island, 31.65667 ° N 81.15167 ° W, 24 Apr 1982 – 29 Apr 1982, Rozen and Favreau, 13, 2♀ ( AMNH). Mississippi: George Co.: Lucedale, 30.925 ° N 88.59 ° W, 24 Apr 1931, H. G. Johnston, 1♀ ( TAMU). Harrison Co.: Biloxi, 30.39583 ° N 88.88528 ° W, 16 Apr 1931, H. G. Johnston, 33, 6♀ Quercus nigra (Fagaceae) , 1♀ ( TAMU). Landon, 30.43778 ° N 89.10278 ° W, 15 Apr 1931, H. G. Johnston, 23 ( TAMU). Long Beach, 30.35028 ° N 89.15278 ° W, 18 Apr 1931, H. G. Johnston, 1♀ ( TAMU). Mississippi City, 30.38167 ° N 89.04389 ° W, 11 Apr 1931, H. G. Johnston, 2♀ ( TAMU). Lauderdale Co.: Meridian, 32.36417 ° N 88.70361 ° W, 28 Apr 1931, H. G. Johnston, 1♀ ( TAMU). Leake Co.: Wiggins, 32.70194 ° N 89.63667 ° W, 5 May 1931, H. G. Johnston, 23 ( TAMU). Oktibbeha Co.: Mississippi State University (A. & M. C.), 33.45472 ° N 88.78861 ° W, 29 May 1931, [collector unknown], 1♀ ( TAMU). Pennsylvania: Schuykill Co.: 5 mi S of Frackville on Rt 81, 40.72248 ° N 76.33146 ° W, 8 Jul 1987, A. G. Wheeler, Jr., Quercus ilicifolia (Fagaceae) , 13, 2♀ ( USNM); 23 Jun 1986, A. G. Wheeler, Jr., Quercus ilicifolia (Fagaceae) , 1♀ ( USNM). South Carolina: Beaufort Co.: Hilton Head Island, 32.19361 ° N 80.73833 ° W, 13 Apr 1974, G. C. Gaumer, 1♀ ( TAMU). Greenville Co.: Greenville, 34.8525 ° N 82.39417 ° W, 20 May 1983, R. S. Peigler, Light Trap, 13, 1♀ ( TAMU); 27 Apr 1985, R. S. Peigler, Light Trap, 13 ( TAMU); 6 May 1986, R. S. Peigler, Light Trap, 13 ( TAMU); 11 May 1985, R. S. Peigler, Light Trap, 1♀ ( TAMU). Oconee Co.: Seneca, 34.68556 ° N 82.95333 ° W, 6 May 1976 – 25 May 1976, R. S. Peigler, Light Trap, 1♀ ( TAMU). Tennessee: Hamilton Co.: unknown, 1 Oct 1943, Turner, 33 ( USNM). Texas: Anderson Co.: 5 mi E Salmon, 31.57275 ° N 95.41288 ° W, 13 Apr 1990, H. R. Burke, E. Riley & C. S. Wolfe, 13 ( TAMU). Angelina Co. : Upland Island Wilderness Area, Graham Creek bottomland, 31.1431 ° N 94.40779 ° W, 9 Apr 1999, W. Godwin, 63, 17♀ ( TAMU); 10 Apr 1999, M. Yoder, Light Trap, 13, 1♀ ( TAMU). Aransas Co.: Goose Island State Park, 28.13528 ° N 96.98722 ° W, 12 Apr 1992, W. F. Chamberlain, 1♀, Light Trap, 83, 3♀ ( TAMU); 22 Apr 1962, W. F. Chamberlain, 13 ( TAMU). Goose Island State Recreation Area, 28.12806 ° N 96.99167 ° W, 12 Apr 1992, W. F. Chamberlain, 23, Light Trap, 33 ( TAMU). Bandera Co. : 12.5 miles W of Medina, 29.79622 ° N 99.45523 ° W, 2 May 1983, J. C. Schaffner, 1♀ ( TAMU). 3 mi E Bandera , 29.72638 ° N 99.02317 ° W, 19 Apr 1984, J.

C. Schaffner, 9♀ ( TAMU). Lost Maples State Natural Area Trails, 29.81522 ° N 99.57505 ° W, 27 Apr 1986, K. Haack, P. Kovarik, 93, 4♀, Light Trap, 1♀ ( TAMU); 28 Apr 1988 – 30 Apr 1988, R. Anderson, 2♀ ( TAMU). Bastrop Co.: Bastrop State Park, 30.11222 ° N 97.26056 ° W, 6 Apr 1959 – 7 Apr 1959, Bottimer, Mason & McAlpine, Light Trap, 2♀ ( CNC). Blanco Co.: 10 mi SE Johnson City, 30.17396 ° N 98.2928 ° W, 29 Apr 1983, J. C. Schaffner, 1♀ ( TAMU). 5 mi E Cypress Mill, 30.3808 ° N 98.16614 ° W, 28 Apr 1971, V. V. Board, 1♀ ( TAMU). Bosque Co.: 3 mi W of Laguna Park, 31.85917 ° N 97.4305 ° W, 21 Apr 1972, J. C. Schaffner, 1♀ ( TAMU). Laguna Park, 31.85917 ° N 97.37944 ° W, 13 Apr 1984, J. B. Woolley, 13 ( TAMU). Brazos Co.: 9 km SSE College Station, 15889 Woodlake Dr., 30.53333 ° N 96.28333 ° E, 8 Apr 2001, J. Oswald, Light Trap, 73, 4♀ ( TAMU). Bryan, 30.67417 ° N 96.36972 ° W, 13 Apr 1965 – 12 Apr 1974, J. C. Schaffner, Light Trap, 23, 1♀ ( JTP). Light Trap, 523, 20♀, 1♀ ( TAMU); 1 May 1965 – 13 May 1970, J. C. Schaffner, Light Trap, 423, 16♀ ( TAMU); 6 Apr 1966 – 8 May 1966, J. C. Schaffner, Light Trap, 63, 4♀ ( TAMU); 27 Apr 1966, J. C. Schaffner, Light Trap, 13 ( TAMU); 13 Apr 1999, Wm. Godwin, 1♀ ( TAMU); 10 Apr 1972, J. C. Schaffner, Light Trap, 23 ( TAMU); 21 Mar 1967, H. R. Burke, 13 ( TAMU); 20 Mar 1967 – 28 Mar 1967, H. R. Burke, 23 ( TAMU); 25 Apr 1970, R. G. Phelps, 23 ( TAMU); 1 Apr 1967, J. C. Schaffner, Light Trap, 13 ( USNM); 11 May 1966, J. C. Schaffner, Light Trap, 1♀ ( USNM); 22 Apr 1965, J. C. Schaffner, Light Trap, 13 ( USNM). Bryan, 30.67417 ° N 96.36972 ° W, 6 Apr 1967, J. C. Schaffner, 1♀ ( TAMU). College Station, 30.62778 ° N 96.33417 ° W, 15 Apr 1978, Henry, Schaffner, and Schuh, Quercus virginiana (Fagaceae) , 13, 1♀ ( AMNH). Quercus virginiana (Fagaceae) , 113, 14♀ ( TAMU); 23 Apr 1996, T. Junek, live oak, 13, 6♀ ( TAMU); 7 Apr 1999, E. G. Riley, 103, 1♀ ( TAMU); 2 Apr 1982, J. C. Schaffner, 53, 5♀ ( TAMU); 15 Apr 1970, J. C. Schaffner, Quercus virginiana (Fagaceae) , 13 ( JTP); 22 Apr 1973, J. C. Schaffner, 13 ( JTP); 14 Apr 1996, T. Junek, live oak, 3♀ ( TAMU); 25 Apr 1970, J. C. Schaffner, Quercus virginiana (Fagaceae) , 23, 3♀ ( TAMU); 1 May 1973, W. E. Clark, 13 ( TAMU); 15 Apr 1965 – 25 Apr 1970, J. C. Schaffner, Light Trap, 13, 5♀ ( TAMU); 20 Apr 1968, J. C. Schaffner, Quercus virginiana (Fagaceae) , 83 ( TAMU); 12 Apr 1964, S. G. Wellso, 13 ( TAMU). College Station, Lick Creek Park, 30.57755 ° N 96.29052 ° W, 6 May 2002, J. C. Schaffner, Light Trap, 1♀ ( TAMU). College Station, Riley Estate, 30.58833 ° N 96.25333 ° E, 10 Apr 2001, E. G. Riley, Light Trap, 53, 1♀ ( TAMU); 12 Apr 2001, E. G. Riley, 13 ( TAMU). Brooks Co.: 7.3 miles S Falfurrias Hwy 281 rest stop, 27.12064 ° N 98.14389 ° W, 8 May 1989, E. G. Riley, 13, 1♀ ( TAMU). Burnet Co.: Inks Lake State Park, 30.73083 ° N 98.38444 ° W, 13 Apr 1985, P. Kovarik, R. Jones, C. Agnew, 13, 2♀ ( TAMU); 28 Apr 1968, J. C. Schaffner, Light Trap, 63, 3♀ ( TAMU); 26 Apr 1969, J. C. Schaffner, Light Trap, 13 ( TAMU); 21 Apr 1986, Kovarik and Haack, 1♀ ( TAMU). Comal Co.: 5 mi W of Sattler on Canyon Lake, 29.84778 ° N 98.25861 ° W, 17 Apr 1983, J. C. Schaffner, 4♀ ( TAMU). Concho Co.: 3 miles E Eden, 31.2161 ° N 99.79436 ° W, 20 Apr 1985, J. C. Schaffner, 33 ( TAMU). Culberson Co.: just N of Guadalupe Mountains National Park, 31.9 ° N 104.86667 ° W, 28 Apr 1978, R. T. Schuh, Quercus sp. (Fagaceae) , 1♀ ( AMNH). Edwards Co.: 14 mi W Rocksprings, 30.01535 ° N 100.43972 ° W, 15 Apr 1985, J. C. Schaffner, 23 ( TAMU). 17 miles NE Rocksprings, 30.18992 ° N 100.00345 ° W, 18 Apr 1984, J. C. Schaffner, 33, 1♀ ( TAMU). 22 mi S of Rocksprings, 29.69751 ° N 100.205 ° W, 30 Apr 1982, J. C. Schaffner, 63 ( TAMU). 27 miles SE Rocksprings, 29.74023 ° N 100.06244 ° W, 18 Apr 1984, J. C. Schaffner, 8♀ ( TAMU). Erath Co.: Stephenville, 32.22056 ° N 98.20194 ° W, 21 Apr 1972, J. C. Schaffner, Light Trap, 13 ( TAMU); 25 Apr 1967, J. C. Schaffner, Light Trap, 13 ( TAMU). Fayette Co.: La Grange, 29.90528 ° N 96.87639 ° W, 7 Apr 1959, W. R. Mason, Light Trap, 13 ( CNC); 7 Apr 1959, J. F. McAlpine, Light Trap, 23 ( CNC). Grimes Co.: Navasota, 30.38778 ° N 96.0875 ° W, 19 Apr 1948, M. Polhemus, 1♀ ( AMNH). Harris Co.: Houston, 29.76306 ° N 95.36306 ° W, 17 Apr 1959 – 19 Apr 1959, E. C. Becker, Light Trap, 213, 5♀ ( CNC). Hays Co.: 1 mi SE Wimberley, 29.98695 ° N 98.08647 ° W, 25 Apr 1982, J. C. Schaffner, 1♀ ( TAMU). 5 mi W of San Marcos, 29.88303 ° N 98.02483 ° W, 19 Apr 1984, J. C. Schaffner, 1♀ ( TAMU). Kendall Co.: Boerne City Park, 29.79444 ° N 98.73167 ° W, 19 Apr 1984, J. C. Schaffner, 13, 5♀ ( TAMU). Kenedy Co.: 20 mi N of Raymondville, 26.77161 ° N 97.78278 ° W, 20 Apr 1974, J. C. Schaffner, 1♀ ( TAMU). 25 mi S Kingsville, 27.15248 ° N 97.85583 ° W, 20 Apr 1974, J. C. Schaffner, 13 ( TAMU). Kenedy Ranch, Jaboncillos Pasture dune area, 26.94213 ° N 97.72257 ° W, 21 Apr 2001, A. Gillogly and J. Schaffner, 33, 49♀, Light Trap, 13, 9♀ ( TAMU); 21 Apr 2001, Raber, Riley and Yoder, Light Trap, 53, 17♀, 83, 19♀ ( TAMU); 20 Apr 2001, J. Oswald and M. Yoder, Light Trap, 53, 14♀ ( TAMU); 21 Apr 2001, W. Godwin, Light Trap, 13, 1♀ ( TAMU); 21 Apr 2001, Raber, Riley, Light Trap, 1♀ ( TAMU). Kenedy Ranch, Jaboncillos Pasture, 0.8 km SW San Pedro Camp, 26.945 ° N 91.66111 ° E, 21 Apr 2001, J. Oswald, Light Trap, 13, 15♀ ( TAMU). Kenedy Ranch, Jaboncillos Pasture, 2 km E Cayo Grande, 27.02027 ° N 97.71166 ° E, 20 Apr 2001, J. Oswald, Light Trap, 13, 5♀ ( TAMU). Kenedy Ranch, Jaboncillos Pasture, San Pedro Camp, 26.95111 ° N 97.65611 ° E, 20 Apr 2001, J. Oswald and M. Yoder, Light Trap, 23, 2♀ ( TAMU); 21 Apr 2001, J. A. Jackman, 1♀ ( TAMU). Kerr Co.: 5 mi N of Kerrville, 30.1198 ° N 99.14 ° W, 18 May 1990, W. F. Chamberlain, Light Trap, 13 ( TAMU). Hunt, 30.07056 ° N 99.3375 ° W, 1 May 1966, J. C. Schaffner, 13 ( TAMU). Kerrville, 30.04722 ° N 99.14 ° W, 10 Apr 1996, W. F. Chamberlain, 1♀ ( TAMU); 15 May 1990, W. F. Chamberlain, 3♀ ( TAMU); 18 Apr 1959, W. R. Mason, 23 ( CNC); 11 Apr 1955, L. J. Bottimer, 23, 1♀ ( CNC); 11 May 1954, L. J. Bottimer, 1♀ ( CNC); 5 May 1968, W. F. Chamberlain, 1♀ ( TAMU); 8 May 1965, W. F. Chamberlain, 1♀ ( TAMU). RNC, Kerrville, 30.04722 ° N 99.14 ° W, 6 Apr 1999, W. F. Chamberlain, 23 ( TAMU). Kimble Co.: 1 mile E Junction, 30.48916 ° N 99.75482 ° W, 17 Apr 1984, J. C. Schaffner, 13, 2♀ ( TAMU). Junction, 30.48917 ° N 99.77167 ° W, 29 Apr 1983, J. C. Schaffner, 1♀ ( TAMU); 17 Apr1984 – 18 Apr 1984, J. C. Schaffner, 83, 6♀ ( TAMU). Leon Co.: 5.5 mi N of Flynn, 31.23034 ° N 96.12417 ° W, 14 May 1997, A. R. Gillogly, 1♀, Light Trap, 43, 3♀ ( TAMU). Llano Co.: 13 miles W of Llano, 30.75898 ° N 98.89432 ° W, 23 Apr 1979, H. Burke, D. Delorme, J. C. Schaffner, 5♀ ( TAMU). Mason Co.: 3 mi NE Streeter, 30.7955 ° N 99.34027 ° W, 17 Apr 1984, J. C. Schaffner, 23 ( TAMU). 8.5 mi S of Mason, Llano River crossing, 30.63672 ° N 99.223 ° W, 13 May 1997, Gilogly and Schaffner, 1♀ ( TAMU). Matagorda Co.: Bay City, 28.9825 ° N 95.96917 ° W, 2 Apr 1976, G. V. Manley, 2♀ ( TAMU). Real Co.: 4 mi n Varice, 29.83316 ° N 99.8384 ° W, 18 Apr 1984, J. C. Schaffner, 2♀ ( TAMU). 5 mi E Leakey, 29.72858 ° N 99.67751 ° W, 1 May 1983, J. C. Schaffner, 13 ( TAMU). 6 mi E Leakey, 29.72857 ° N 99.66079 ° W, 18 Apr 1984, J. C. Schaffner, 13, 3♀ ( TAMU). San Patricio Co.: Aransas Pass, 27.90917 ° N 97.14972 ° W, 11 Apr 1939, G. Willett, 13, 1♀ ( LACM). Lake Corpus Christi State Park, 28.06167 ° N 97.87639 ° W, 16 May 1965, M. H. Sweet, 13, 1♀ ( TAMU). Mathis, 28.09417 ° N 97.82778 ° W, 4 Jun 1964, H. R. Burke, 1♀ ( TAMU). Welder Wildlife Refuge, near Sinton, 28.03639 ° N 97.50889 ° W, 19 Apr 1983, T. J. Henry and A. G. Wheeler, Jr., Celtis laevigata (Ulmaceae) , 1♀ Prosopis glandulosa (Fabaceae) , 43, 1♀ ( USNM); 18 Apr 1989 – 19 Apr 1989, R. Anderson, 53, 5♀ ( TAMU). San Saba Co.: 5 miles SE of Bend, 31.0481 ° N 98.45145 ° W, 21 Apr 1997, W. F. Chamberlain, 13, 3♀ ( TAMU). Shackelford Co.: Ford Griffin State Park, 32.92277 ° N 99.22472 ° E, 10 May 2002, E. G. Riley, 1♀ ( TAMU). Smith Co.: Tyler State Park, 32.35111 ° N 95.30083 ° W, 9 May 1988, R. S. Anderson, Light Trap, 13 ( TAMU). Somervell Co.: 10 mi W Glen Rose, 32.23432 ° N 97.92657 ° W, 4 May 1973, J. C. Schaffner, 1♀ ( TAMU). Sutton Co.: 16 mi W of Sonora, 30.56667 ° N 100.91169 ° W, 11 May 1997, Gillogly, Schaffner, 13, 2♀ ( TAMU). 17 miles E of Sonora, 30.56635 ° N 100.35645 ° W, 27 Apr 1985, W. F. Chamberlain, 33 ( TAMU). Sonora, 30.56667 ° N 100.64306 ° W, 29 Apr 1932, S. E. Jones, 223, 8♀ ( TAMU). Travis Co.: Zilker Park, 30.26722 ° N 97.76972 ° W, 3 Apr 1985, P. W. Kovarik, 13, 2♀ ( TAMU). Vicinity of Long Hollow Creek, 30.44781 ° N 97.88212 ° W, 23 Apr 1994, M. Quinn, E. Riley, R. Wharton, Quercus virginiana (Fagaceae) , 13, 4♀ ( TAMU); 23 Apr 1993, Alexander, Quinn, Riley, Wharton et al., Ulmus crassifolia (Ulmaceae) , 13 Quercus buckleyi (Fagaceae) , 13, 1♀ Quercus virginiana (Fagaceae) , 53, 7♀ ( TAMU); 22 Apr 1993, [collector unknown], 1♀ ( TAMU); 8 May 1993, Alexander, Quinn, Riley, Wharton et al., Quercus virginiana (Fagaceae) , 2♀ ( TAMU). Uvalde Co.: 1.3 miles NW Uvalde, 29.22278 ° N 99.80112 ° W, 15 Apr 1985, J. C. Schaffner, 23 ( TAMU). 16.4 miles NW Uvalde (on Nueces River), 29.37192 ° N 99.98607 ° W, 15 Apr 1985, J. C. Schaffner, 13, 6♀ ( TAMU). 29 miles NW Uvalde, 29.52295 ° N 100.03118 ° W, 15 Apr 1985, J. C. Schaffner, 53, 2♀ ( TAMU). Val Verde Co. : 2.7 mi W of Comstock, 29.68417 ° N 101.21798 ° W, 16 Apr 1985, J. C. Schaffner, 13 ( TAMU). Del Rio, 29.3625 ° N 100.89639 ° W, 29 Apr 1959, W. R. Mason, Light Trap, 13 ( CNC). Williamson Co.: Taylor, 30.57056 ° N 97.40917 ° W, 26 Apr 1968, J. E. Hafernik, 23 ( TAMU). Zavala Co.: Crystal City, 28.67722 ° N 99.82778 ° W, 24 Apr 1978, T. J. Henry and R. T. Schuh, Quercus sp. (Fagaceae) , 1♀ ( TAMU).