Timandra accumulata, Cui, Le, Xue, Dayong & Jiang, Nan, 2019

Cui, Le, Xue, Dayong & Jiang, Nan, 2019, A review of Timandra Duponchel, 1829 from China, with description of seven new species (Lepidoptera, Geometridae), ZooKeys 829, pp. 43-74 : 43

Timandra accumulata sp. n. Figs 11, 29, 40, 52, 68

Description.

Head. Antennae bipectinate in basal five-sixths in male, filiform in female; dorsal surface of shaft pale yellowish-brown with brown scales except in basal part. Frons blackish-brown, mixed with yellowish-white on ventral side, forming a rounded protrusion (Fig. 40). Labial palpi yellowish-brown, not extending beyond frons. Vertex yellowish-white. Thorax. Patagia deep brown. Tegulae and thorax greyish-brown. Hind tibia with two pairs of spurs in male. Forewing length: male 16-18 mm. Forewing with acute apex; outer margin almost straight; apex of hindwing rounded; outer margin protruding on vein M3. Wing colour pale yellowish-brown densely scaled with greyish-brown spots. Forewing with indistinct antemedial line angled at cell; discal spot greyish-brown and short bar-like; medial line deep brown and oblique, raising from apex and extending to middle part of terminal margin, gradually broadening towards termen; postmedial line greyish-brown, overlapping with medial line at apex, separated from it before R5, slightly protruding; terminal line greyish-brown and narrow; fringes yellowish-brown. Hindwing with medial line deep brown and straight; postmedial line greyish-brown, slightly curved outwards in middle part; terminal line and fringes similar to those of forewing. Underside with deep greyish-brown spots, discal spot stronger and longer than upperside; postmedial line more distinct than that on upperside.

Male genitalia. Uncus broad. Tegumen narrow. Socii straight, narrow and rod-like, slightly extending beyond tip of uncus. Costa of valva thickened and slightly curved at terminal half, quadrate apically; valvula forming a rounded protrusion and with bristles at apex, arm between valvula and sacculus short and digitiform with teeth on ventral margin, a small process present on base of costa, covered with bristles; sacculus slightly shorter than valvula and asymmetrical, right side slightly shorter than left side, apex of right side acute and rounded on left side; densely covered with long bristles terminally. Juxta with broad base, tapered towards terminal part. Saccus broad and rounded at apex. Aedeagus short, a long sclerotized band present on vesica.

Female genitalia. Seventh sternite sclerotized, bifurcate at middle and forming two rounded lateral processes on posterior margin. Colliculum long. Ductus bursae short and membranous. Corpus bursae oval; signum present as a triangular pouch directed towards anterior part, with a longitudinal ridge running from apex.

Diagnosis.

The species is characterized by the very dense greyish-brown spots on the wings, which is similar to T. rectistrigaria (Eversmann, 1851), but the male and female genitalia of these two species are quite different ( Hausmann 2004: fig. 177). The male genitalia are characterized by the following features: the uncus is stout; the socii are narrow and straight; the costa of the valvula is slightly curved, the arm between the valvula and the sacculus is short; the right side of the sacculus is shorter than the left side and acute apically; the left side of the sacculus is rounded apically. The female genitalia are similar to those of T. recompta , but the seventh sternite of T. accumulata is broader, the colliculum is longer in T. accumulata , and the appendix bursae of T. accumulata is absent.

Type material.

Holotype, ♂, CHINA: Yunnan (IZCAS): Lijiang, Yulong Shan, 10.VII.1962, coll. Song Shimei. Paratypes: 2♂, same as holotype, 21.VII.1962, coll. Song Shimei. Yunnan (ZFMK): 73♂13♀, Yunnan, Li-kiang, 6. VI– 31.VIII.1934, 3-24.VII.1935, 12.VI.1935, 20.VI.1935, 21.IX.1935, 3.IX.1935, coll. H. Höne.

Distribution.

China (Yunnan).

Etymology.

The species is named based on the Latin accumulatus, which refers to the dense pattern of brown spots accumulated on the wing surface.