Harpalus somnulentus Dejean, 1829

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 705-707

Harpalus somnulentus Dejean, 1829

Harpalus somnulentus Dejean, 1829: 333. Type locality: «détroit de Norfolk [= Norfolk Sound, Baranof Island, Alaska] sur la côte nord-ouest de l’Amérique septentrionale" (original citation). Lectotype (♂), designated by Noonan (1991: 190), in MHNP.

Harpalus pleuriticus Kirby, 1837: 41. Type locality: "Lat. 54° [= along North Saskatchewan River]" (original citation), restricted to "Edmonton, Al[ber]ta" by Lindroth (1968: 792). Lectotype (♂), designated by Noonan (1991: 190), in BMNH. Synonymy established by Noonan (1991: 190).

Harpalus fallax LeConte, 1859c: 2. Type locality: "Santa Fé [Santa Fe County, New Mexico]" (original citation). Lectotype (♂), designated by Noonan (1991: 190), in MCZ [# 5889]. Synonymy established by Noonan (1991: 190).

Harpalus stupidus LeConte, 1859c: 3. Type locality: "route to Fort Bridger" (original citation). Holotype [by monotypy] (♀) in MCZ [# 5906]. Synonymy established by Noonan (1991: 191).

Harpalus viduus LeConte, 1859c: 3. Type locality: "New Jersey" (original citation). Syntype(s) presumably lost. Synonymy established, under the name Harpalus fallax LeConte, by Lindroth (1968: 794).

Harpalus carbonatus LeConte, 1860: 319. Type locality: «Saskatchewan» (original citation). Lectotype (♀), designated by Noonan (1991: 191), in MCZ [# 5896]. Synonymy established by Noonan (1991: 191).

Harpalus placidus Casey, 1884b: 10 [primary homonym of Harpalus placidus Gyllenhal, 1827]. Type locality: "N[ew] J[ersey]" (holotype label). Holotype [by monotypy] (♂) in USNM [# 47807]. Synonymy established, under the name Harpalus fallax LeConte, by Horn (1885b: 109), confirmed by Noonan (1991: 191). Note. "Willets Point, Long Island" was the locality cited in the original description.

Harpalus recisus Casey, 1914: 93. Type locality: "New Jersey" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47791]. Synonymy established, under the name Harpalus fallax LeConte, by Lindroth (1968: 794).

Harpalus futilis Casey, 1914: 97. Type locality: "Lake Tahoe [Placer County], California" (original citation). Holotype [by monotypy] (♂) in USNM [# 47803]. Synonymy established by Noonan (1991: 191).

Harpalus intactus Casey, 1914: 97. Type locality: «California» (original citation). Holotype [by monotypy] (♀) in USNM [# 47795]. Synonymy established by Noonan (1991: 192).

Harpalus aequabilis Casey, 1914: 100. Type locality: "Buena Vista [Chaffee County], Colorado" (original citation). Lectotype (♀), designated by Lindroth (1975: 139), in USNM [# 47779]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Lindroth (1954b: 142).

Harpalus lascivus Casey, 1914: 100. Type locality: "British Columbia" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47800]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Hatch (1953: 169), confirmed by Noonan (1991: 192).

Harpalus pumilio Casey, 1914: 100 [primary homonym of Harpalus pumilio Dejean, 1829]. Type locality: "Bayfield [Bayfield County], Wisconsin" (original citation). Lectotype (♀), designated by Lindroth (1975: 139), in USNM [# 47801]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Lindroth (1954b: 142).

Harpalus perspicuus Casey, 1914: 101. Type locality: "Boulder Co[unty], Colorado" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47797]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Lindroth (1954b: 142).

Harpalus lividulus Casey, 1914: 101. Type locality: "Bayfield [Bayfield County], Wisconsin" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47804]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Lindroth (1954b: 142).

Harpalus pellax Casey, 1914: 105. Type locality: "Yellowstone National Park" (original citation). Lectotype (♂), designated by Noonan (1991: 190), in USNM [# 47812]. Synonymy established by Noonan (1991: 192).

Harpalus peritus Casey, 1914: 107. Type locality: "Reno [Washoe County], Nevada" (original citation). Lectotype (♂), designated by Noonan (1991: 190), in USNM [# 47814]. Synonymy established by Noonan (1991: 192).

Harpalus celox Casey, 1914: 107 (as celax, typographic error, see Casey 1914: 383). Type locality: "Clackamas Co[unty], Oregon" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47816]. Synonymy established by Noonan (1991: 191).

Harpalus uteanus Casey, 1914: 118. Type locality: "Provo [Utah County], Utah" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47835]. Synonymy established by Noonan (1991: 191).

Harpalus oppositus Casey, 1914: 125. Type locality: "Siskiyou Co[unty], California" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47860]. Synonymy established, under the name Harpalus celox Casey, by Hatch (1953: 169), confirmed by Lindroth (1968: 789).

Harpalus amiculus Casey, 1924: 102. Type locality: "Lake George [Warren County], New York" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47792]. Synonymy established, under the name Harpalus fallax LeConte, by Lindroth (1968: 794).

Harpalus contactus Casey, 1924: 106. Type locality: "Cedar City [Iron County], Utah" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47818]. Synonymy established by Lindroth (1968: 789).

Harpalus uintanus Casey, 1924: 106. Type locality: "Cedar City [Iron County], Utah" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47820]. Synonymy established, under the name Harpalus intactus Casey, by Lindroth (1968: 789).

Harpalus blanditus Casey, 1924: 108. Type locality: "Terrace, British Columbia" (original citation). Lectotype (♂), designated by Lindroth (1975: 139), in USNM [# 47802]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Hatch (1953: 169), confirmed by Noonan (1991: 193).

Harpalus nivalis Casey, 1924: 109. Type locality: "Redvers, Sask[atchewan]" (holotype label). Holotype [by monotypy] (♂) in USNM [# 47805]. Synonymy established, under the name Harpalus pleuriticus Kirby, by Lindroth (1954b: 142).

Harpalus pacificus Fall, 1926a: 136. Type locality: "Ketchikan, Alaska" (original citation). Holotype (♂) in MCZ [# 23876]. Synonymy established by Lindroth (1968: 789).

Harpalus rewolinskii Noonan, 1991: 214. Type locality: "2.3 mi[les] S[outh] Oyen, Alberta" (original citation). Holotype (♂) in CNC [# 23531]. Synonymy established by Ball and Bousquet (2000: 99).

Distribution.

This species is transcontinental in the north ranging from Newfoundland to the Kenai Peninsula in Alaska, south to southern California, central Arizona and New Mexico in the Rocky Mountains, Nebraska, and northern Virginia [see Noonan 1991: Figs 297-309]. The records from central Alabama (see Noonan 1991: Figs 302 and 304), “Arkansas” (Bousquet and Larochelle 1993: 237) and southern Kansas (Snow 1903: 194, as Harpalus fallax ) need confirmation.

Records.

FRA: PM CAN: AB, BC (QCI, VCI), LB, MB, NB, NF, NS (CBI), NT, ON, PE, QC, SK, YT USA: AK, AZ, CA, CO, CT, DE, IA, ID, IL, IN, MA, MD, ME, MI, MN, MO, MT, ND, NE, NH, NJ, NM, NV, NY, OH, OR, PA, RI, SD, UT, VA, VT, WA, WI, WV, WY [AL, AR, KS]

Note.

Noonan (1991: 196-207) recognized six morphs within this species, the pleuriticus, fallax, “dark,” uteanus, somnulentus, and carbonatus morphs, which Lindroth (1968) treated as distinct species, except for the “dark” one. Noonan went on to report numerous intergradation zones between all morphs.