Cionocoleus longicapitis, Soriano & Delclòs, 2006

Soriano, Carmen & Delclòs, Xavier, 2006, New cupedid beetles from the Lower Cretaceous of Spain and the palaeogeography of the family, Acta Palaeontologica Polonica 51 (1), pp. 185-200 : 187-188

Cionocoleus longicapitis sp. nov.

Figs. 2C , 3A .

Type locality: Quarry near Rubies , La Pedrera de Meià fossil site, Sta. Maria de Meià , Sierra del Montsec, Lleida Province, Spain .

Stratigraphic horizon: Calcaires lithographiques à Plantes et Vértébres de la Pedrera de Rubies Fm., Lower Cretaceous, Barremian ( Martín−Closas and López−Morón 1995).

Derivation of the name: After the long head of this species.

Material.— Holotype IEI LP 0164 G, only the part.

Diagnosis.—Distinct from Cionocoleus magicus Ren, 1995 in elongate elytra, but without acuminate apex like in C. sibirica Ponomarenko, 2000 . From C. ommamimus Ponomarenko, 1997 differs in the ratio between abdominal ventrites and length of the head.

Description.—Measurements: length 19.0 mm; width 7.0 mm; elytra length 13.0 mm. Body rather large and elongate, general shape almost subcylindrical. Head transverse, about two times as wide as long without mandibles; genae as long as eyes, temples rounded and two times shorter than eyes. Mandibles with teeth in vertical plane. Antennae moniliform, antennomeres rounded somewhat wider at posterior edge. Eyes comparatively large and convex. Neck−like constriction not distinct. Pronotum elongate and narrowed towards base and apex, about 1.3 times as long as wide, somewhat wider than head, lateral margins serrate, anterior and posterior angles rounded. Procoxae rounded and contiguous. Mesosternum transverse, about 2.5 times as wide as long. Metasternum 1.3 times as wide as long at its posterior margin, with well expressed transverse and longitudinal sutures. Abdomen with 5 visible ventrites, last ventrite about 1.2 times longer than previous one. Elytra about 3.3 times as long as wide, and almost parallel−sided, epipleura comparatively narrow. Cells on elytra not visible. Profemora somewhat widened posteriorly at its medial length, almost as long as protibiae. Body densely covered with tubercles, larger on head and pronotum, and quite smaller in abdomen.

Discussion.—This beetle is similar to beetles of the genus Omma Ponomarenko, 1969 in almost all characters, except for smooth elytra. Cionocoleus was described by isolated elytra ( Ren 1995), and Ponomarenko (1997) placed this genus into Cupedidae , but because of the lack of preservation of the fore coxae was not possible to assign it to any subfamily. The new material from the fossil sites of Spain allows to place this genera in the subfamily Ommatinae , tribe Ommatini , by the presence contiguous fore coxae, not divided by any process.

Figures

Fig. 2. Barremian (Early Cretaceous) ommatins Tetraphalerus, Cionocoleus (both Ommatini), Brochocoleus (Brochocoleini), and Zygadenia (Notocupedini) from Spain. A. Tetraphalerus ponomarenkoi sp. nov. Holotype MCCM LH 78a, part, Las Hoyas, Cuenca Province. B. Tetraphalerus penalveri sp. nov. Holotype DEPGM LP 0058b, counterpart, La Pedrera, Lleida Province. C. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, Lleida Province. D. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, Lleida Province. E. Zygadenia viridis sp. nov. Holotype MCCM LH 1805a, part, Las Hoyas, Cuenca Province. F. Zygadenia oculata sp. nov. Holotype IEI LC 3675a, part, La Cabrúa, Lleida Province. Scale bars 2 mm.

Fig. 3. Camera lucida drawings of Barremian (Early Cretaceous) ommatin Cionocoleus and brochocoleinin Brochocoleus from Lleida Province of Spain. A. Cionocoleus longicapitis sp. nov. Holotype IEI LP 0164G, La Pedrera, dorsal (A1) and ventral (A2) aspects. B. Brochocoleus indibili sp. nov. Holotype IEI LC 4636a, part, La Cabrúa, dorsal (B1) and ventral (B2) aspects. Scale bars 2 mm.