Calolydella nigripalpis Fleming & Wood

Fleming, AJ, Wood, D. Monty, Smith, M. Alex, Hallwachs, Winnie & Janzen, Daniel H, 2018, Revision of the Mesoamerican species of Calolydella Townsend (Diptera: Tachinidae) and description of twenty-three new species reared from caterpillars in Area de Conservacion Guanacaste, northwestern Costa Rica, Biodiversity Data Journal 6, pp. 11223-11223 : 11223

Calolydella nigripalpis Fleming & Wood sp. n.

Materials

Type status: Holotype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017812 ; recordedBy: D.H. Janzen, W. Hallwachs & Carolina Cano; individualID: DHJPAR0017812; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-1172, BOLD:ABY2592, ASTAR523-07; Taxon: scientificName: Calolydellanigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Puente Palma; verbatimElevation: 460; verbatimLatitude: 10.9163; verbatimLongitude: -85.3787; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 04-Apr-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0040836 ; recordedBy: D.H. Janzen, W. Hallwachs & Ricardo Calero; individualID: DHJPAR0040836; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 10-SRNP-73126, BOLD:ABY2592, ASHYF751-11; Taxon: scientificName: Calolydellanigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Guanacaste; county: Sector Pitilla; locality: Area de Conservacion Guanacaste ; verbatimLocality: Calma; verbatimElevation: 412; verbatimLatitude: 11.0099; verbatimLongitude: -85.3921; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 20-Nov-2010; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0052029 ; recordedBy: D.H. Janzen, W. Hallwachs & Edwin Apu; individualID: DHJPAR0052029; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 13-SRNP-69658, BOLD:ABY2592, ASHYH1141-13; Taxon: scientificName: Calolydellanigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Jacobo; verbatimElevation: 461; verbatimLatitude: 10.9408; verbatimLongitude: -85.3177; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 26-May-2013; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0017811 ; recordedBy: D.H. Janzen, W. Hallwachs & Yessenia Mendoza; individualID: DHJPAR0017811; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 05-SRNP-4251, BOLD:ABY2592, ASTAR522-07; Taxon: scientificName: Calolydellanigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector San Cristobal; locality: Area de Conservacion Guanacaste ; verbatimLocality: Potrero Argentina; verbatimElevation: 520; verbatimLatitude: 10.8902; verbatimLongitude: -85.388; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 20-Aug-2005; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054198 ; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0054198; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 14-SRNP-75117, BOLD:ABY2592, ASHYD3366-14; Taxon: scientificName: Calolydellanigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 29-Jan-2014; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen Type status: Paratype. Occurrence: occurrenceDetails: http://janzen.sas.upenn.edu; catalogNumber: DHJPAR0054195 ; recordedBy: D.H. Janzen, W. Hallwachs & Cirilo Umana; individualID: DHJPAR0054195; individualCount: 1; sex: M; lifeStage: adult; preparations: pinned; otherCatalogNumbers: 14-SRNP-75120, BOLD:ABY2592, ASHYD3363-14; Taxon: scientificName: Calolydellanigripalpis; phylum: Arthropoda; class: Insecta; order: Diptera; family: Tachinidae; genus: Calolydella; specificEpithet: nigripalpis; scientificNameAuthorship: Fleming & Wood, 2016; Location: continent: Central America; country: Costa Rica; countryCode: CR; stateProvince: Alajuela; county: Sector Rincon Rain Forest; locality: Area de Conservacion Guanacaste ; verbatimLocality: Estacion Llanura; verbatimElevation: 135; verbatimLatitude: 10.9333; verbatimLongitude: -85.2533; verbatimCoordinateSystem: Decimal; Identification: identifiedBy: AJ Fleming; dateIdentified: 2016; Event: samplingProtocol: Reared from the larva of the geometrid moth, Semaeopus Janzen08; verbatimEventDate: 01-Feb-2014; Record Level: language: en; institutionCode: CNC; collectionCode: Insects; basisOfRecord: Pinned Specimen

Description

Male (Fig. 24a, b, c). Length: 5-6mm. Head (Fig. 24b): frontal setae extending to base of postpedicel; fronto-orbital plate gold, sparsely setulose throughout; parafacial silver throughout; palps black. Thorax (Fig. 24a, c): gold on both dorsal and lateral surfaces; four almost indistinct thoracic vittae barely visible except under certain angles of light; postpronotum with three setae; 3:3 acrostichal setae; 3:3 dorsocentral setae; 2:3 intra-alar setae; 2:3 supra-alar setae; three katepisternal setae; anatergite bare; scutellar discal setae situated as wide apart as subapical scutellar setae. Wing vein R4+5 with 6-7 setulae dorsally, reaching from the base almost to crossvein R-M. Abdomen (Fig. 24a): ground color black, with uninterrupted transverse marginal pollinose bands; pollinosity gold dorsally, silver ventrally; base of ST1+2 black lateroventrally; T3 with one pair of median marginal setae and two pairs of discal setae; T4 with a row of marginal setae and two pairs of discal setae. Terminalia (Fig. 25): sternite 5 (Fig. 25c) with two small lobes and a wide U-shaped median cleft, 0.62X the length of the sternite from lobe to apex; inner margin covered by dense pollinosity, appearing darker than surrounding cuticle; with 4-6 short setae, of varying lengths. Cerci (Fig. 25b), in dorsal view, separated by a narrow gap, divergent at apex; each cercus sharply tapered, 2.2X broader at base than at apex, digitiform when viewed dorsally; cercus with a very slight downward bend at tip when viewed laterally; setose along its entire length. Surstylus (Fig. 25a) digitiform, with a strong apical downward curve giving a hooked tip, appearing 0.2X longer than cercus when viewed laterally; with short setae along entire length; tip of surstylus appearing curved inwards when viewed dorsally.

Female: not known at this time.

Diagnosis

Calolydella nigripalpis can be distinguished from all other species of Calolydella by the following combination of characters: frontal setae not extending beyond base of postpedicel, palps black in male, four barely visible thoracic vittae, anatergite bare, and abdominal ground color black.

Etymology

The specific epithet is derived from the Latin adjective "nigrum", meaning black, and "palpus", meaning palp, in reference to the black palpi of males of this species.

Distribution

Costa Rica, ACG, Alajuela and Guanacaste provinces, 135-520m.

Ecology

Calolydella nigripalpis has been reared six times from Semaeopus Janzen08 ( Lepidoptera : Geometridae ), in rain forest.