Staminodeus forcipis, Nico M. Franz, 2001

Nico M. Franz, 2001, Description And Phylogeny Of Staminodeus, A New Genus Of Derelomini (Coleoptera: Curculionidae) Associated With Cyclanthaceae, The Coleopterists Bulletin 55 (4), pp. 411-432 : 425-426

Staminodeus forcipis , new species

Diagnosis. Staminodeus forcipis is distinguished from S. bispinosus by the fairly large frontal spinule and the distinctly arcuate, apically dentate profemur in males. See diagnosis of S. bispinosus for females.

Male. Length 2.1 mm, width 1.0 mm, l/ w 2.1. Rostrum 0.5 mm, r/p 0.8. Frons with 1 apically directed, arcuate spinule, nearly 2/5 as long as diameter of eye, anteriorly with long, fine, erect setae. Pronotum l/ w 0.9, convex. Prosternum longer than mesosternum. Profemur ( Fig. 3e ) (distinctly) arcuate, f/p 1.2, width similar throughout, each ventrolateral margin with row of denticuli/dentes, larger towards apex; protibia ( Fig. 3e ) slightly arcuate, t/f 1.0, anteroventral margin with row of 8–10 apically directed, arcuate dentes, apically mucronate, promucron nearly 2/3 as long as protarsal claw; mesotibia apically inermous. Elytron l/ w 1.4. Median lobe ( Fig. 4e ) l/ w 2.9 (N 1), laterobasal sclerites apically heavily sclerotized, broadened, denticulate, median sclerite basally projected, apically acuminate, glabrate, apical rami distinctly undulate in central region, basal 3/4 of median lobe denticulate, aedeagal apodemes shorter than median lobe.

Female. Length 1.9–2.0 mm, width 1.0 mm, l/ w 1.9 –2.0. Rostrum 0.5 mm, r/p 1.0. Frontal spine nearly 3/5 as long as diameter of eye. Pronotum l/ w 0.8 –0.9. F/p

1.1–1.2; t/f 0.9–1.0. Elytron l/ w 1.4 (N 2). Spermatheca ( Fig. 5e ) deflexed nearly 90, subapically undulate, apex slightly separated, rotundate.

Type Information. Male holotype ‘‘ COSTA RICA, S. J., Zurqui de Moravia , 1,600 m, XI­1995, malaise trap, Hanson & Godoy’ ’ ( CWOB) ; female allotype ‘‘ COSTA RICA, S. Jose, Zurqui de Moravia , 1,600 m, malaise trap, III­1992, P. Hanson’ ’ ( CWOB) ; female paratype ‘‘ COSTA RICA, S. Jose, Zurqui de Moravia , 1,600 m, malaise trap, II­1993, P. Hanson’ ’ ( MUCR, 1: teneral).

Etymology. Named for the dentate profemur and protibia in males which resemble a pair of tongs— forceps signifying ‘‘nippers, tongs’’ ( Brown 1956).

Distribution. Staminodeus forcipis has been collected in the Cordillera de Tilarán of Costa Rica, Provincia Puntarenas, Monteverde; and in the Central Valley of Costa Rica, Provincia San Jose ´, Zurquí de Moravia ( Fig. 6 ).

Natural History. Unknown.

Figures

Fig. 3. Staminodeus, male, right profemur and protibia, anterior view, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Scale bar 0.1 mm.

Fig. 4. Staminodeus, male, aedeagus, ventral and lateral view, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Illustrations are interpretative: denticulation and setation are omitted. Scale bar 0.1 mm.

Fig. 5. Staminodeus, female, spermatheca and gland reservoir, a) S. inermis; b) S. curvitibialis; c) S. denticulatus; d) S. bispinosus; e) S. forcipis; f) S. dilatatus; g) S. vectoris. Scale bar 0.05 mm.

Fig. 6. Distribution of Staminodeus in Costa Rica, Panama, and Venezuela, S. bispinosus; S. curvitibialis (distributional record for Colombia, Provincia Nariño, La Planada is omitted); S. denticulatus; S. dilatatus; S. forcipis; S. inermis; S. vectoris.