Drasteria cailino obscura ( Staudinger, 1901 )

Yu, Alexey & Korb, Stanislav K., 2019, A revision of the genus Drasteria of Central Asia and Kazakhstan with special attention to the adjacent areas (Lepidoptera: Erebidae), Zootaxa 4673 (1), pp. 1-104 : 9-10

Drasteria cailino obscura ( Staudinger, 1901)

( Figs. 1–3 (imago), 208, 209 (male genitalia))

Staudinger in Staudinger et Rebel, 1901: 243 ( Leucanitis Cailino Lef. v. Obscura). Type locality: “Ross. mer. or. (Narün), Tarb. et Ala Tau (trans.)” (by original description); “Narün” (by the lectotype designation, designated here) [“nordöstlich von Astrachan zwischen Wolga- und Ural-Fluss, … etwa 15 deutsche Meilen ostwärts von der Wolga” ( Staudinger, 1879: 315)].

Type material: Lectotype male (designated here in accordance with ICZN Art. 74) ( Figs. 1–3 ), labelled: “Narün | Henke” (green paper, handwritten), “Origin.” (pink paper, printed), “ex coll. 1/6 | Staudinger” (white paper, printed with handwritten numbers), “ LECTOTYPUS ♂ | obscura | Stgr. | S.K. Korb des. 2016” (red paper, printed with handwritten taxon name and author) ( ZMHU) . Paralectotypes 3 males, 2 females ( ZMHU) .

Butler, 1892: 315 [ Asia Minor, Caucasus, Schuscha]; Culot, 1913: 184 [la Russie méridionale]; Aibasov, 1974: 122 [Shevchenko, Mangyshlaksky Karatau: Syir-Su dwell]; Kumakov, Korshunov, 1979: 153 [4 th Dacha, Saratov]; Aibasov, Zhdanko, 1982: 30 [Aksu in Zelinograd Prov.]; Ishkov, Sukhareva, 1986: 124 [Aksu-Dzhabagly Nature Reserve: Novonikolaevka, Dzhabagly, Aksu river canyon, Kshi-Kaindy, Ulken-Kaindy, Aksay]; Poole, 1989: 329; Anikin et al., 2000: 332 [Saratov]; Hacker, Miatleuski, 2001: 814 [Bisen]; Nupponen, Fibiger, 2002: 135 [Arkaim, Bajmak, Kidriasovo, Kuvandyk, Moskovo, Sakmara river, Verbljushka]; Poltavsky, Ilyina, 2002: 17 [Gunib, Tshervlennie Buruni]; Poltavsky, Nekrasov, 2002: 26 [Rostov-on-Don Region, Daghestan Republic]; Goater et al., 2003: 79 [South Europe, in Portugal, Spain, south France, Italy, Sicily and the Balkan countries. Recorded also from South Russia: Saratov District and Kazakhstan: Bisen. Outside Europe, in Middle East, Asia Minor, and Central Asia]; Poltavsky, Ilyina, 2003: 171 [Leninaul, Tchervlennie Buruni]; Poltavsky et al., 2010: 93 [Botkul]; Kononenko, 2010: 89 [ Kazakhstan]; Gorbunov, 2011: 56 [steppes and northern deserts of West Kazakhstan]; Lehmann & Zahiri, 2011: 150 [ Iran: Gate Dehh, Yush]; Volynkin, 2012: 27 [ Kazakhstan part of Altai]; Poltavsky, 2016: 19 [Rostov-on-Don Province]; Shovkoon, Trofimova, 2016: 606 [Aktobe Prov., near Kompressorniy vill. N 48°30’; E 59°19’; Aktobe Prov., Mugodzhary Mts., N 48°26’; E 58°24’; Aktobe Prov., Bolshie Barsuki sands, N 46°58’; E 59°13’; Aktobe Prov., Shagyrai Plateau, N 46°44’; E 57°30’; Kyzyl-Orda Prov., Karatup penin., Aral lake, N 46°20’; E 59°43’]; Titov et al., 2017 a: 148 [Pavlodar region, vic. of Pavlodarskoe vill., 52°22.084’N 76°52.161’E; Pavlodar region, Tuz lake, 51°17’58.24”N 78°37’57.06”E; Pavlodar region, Bayanaul Mts., Birzhankol lake, 50°49’9.97”N 75°20’52.44”E; Pavlodar region, Bayanaul Mts., Kurkeli natural landmark, 50°44’34.00”N 75°38’16.80”E; Pavlodar region, Bayanaul Mts., vic. of Kempirtas Mt., 50°51’24.65”N 75°34’37.21”E; Pavlodar region, Bayanaul Mts., vic. of Toraighyr lake, 50°51’54.01”N 75°40’21.29”E; Pavlodar region, Bayanaul Mts., Kirgichi natural landmark, 50°50’26.05”N 75°44’15.03”E; Pavlodar region, Zhartas natural landmark, 51°38’7.49”N 74°39’54.86”E].

Habitat ( Fig. 299 ). Steppes, deserts and semideserts. Elevation: 200–2700 m.

Distribution. Astrakhan, Saratov, Orenburg and Volgograd Prov., Krasnodarsky Kray, Daghestan ( Russia), Ukraine, North and Central Kazakhstan, Armenia, Azerbaijan, North-West Iran, North-West Turkey.

Material examined (in total: 221 specimens). Armenia: 1 ♂, 1.08.1934, Alagez , southern slope, 8500’, [leg. M. Ryabov] ( ZISP) ; 1 ♂, 18.07.1936, Alagez vic., Inaklu river , [leg. M. Ryabov] ( ZISP) ; 1 ♂, 8.07.1937, Arpa river , [leg. M. Ryabov] ( ZISP) ; 1 ♀, 16.07.1939, Gedzhanan , h=7000–8000’, meadow, [leg. M. Ryabov] ( ZISP) ; 1 ♀, 28.05.1960, Erevan, leg. Vardikian ( ZISP) ; 1 ♂, 15.07.1963, Daralagez, Ger-Ger , [leg. E. Milyanovsky] ( ZISP) ; 1 ♂, 18.06– 7.07.1996, Gegamsky range, Vedy river, right bank, Khosrov reserve , h= 2000 m, leg. A. Danchenko, ex coll. А. Nekrasov, slide ffi2748 Nekrasov ( ZISP) ; 1 ♂, 20.06.2002, Gegamsky range, Khosrov res., leg. M. Ka- lashian, ex coll. А. Nekrasov ( ZISP) ; 3 ♂, 18.07.1936, Aragaz Mt., Antarut ( Inaklyu ), leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 10.07.1965, Aragaz Mt., Kara-Gel’ lake , 3300 m, leg. A.V. Zvetaev ( ZMMU) ; 1 ♂, 4.06.1970, Nakhichevan, Daralagezsky Mts., Buzgov , 1400 m, leg. A.V. Zvetaev ( ZMMU) . Azerbaijan: 1 ♂, Helendorf , ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, Ordubad, leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 4 ♀, 15.06.1881, 7.05– 15.06.1883, Ordub [ad], leg. Chr [istoph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 9.07.1937, Sultanbek , [leg. М. Ryabov] ( ZISP) ; 1 ♀, 20.08.1939, Kupudzhik , upper forest zone, h=8000’, [leg. М. Ryabov] ( ZISP) ; 1 ♂, 14.06.1967, Ordubad, Pazmara , h=1800 м, leg. R. Effendi ( ZISP) ; 1 ♂, 22.07.1986, Nakh- ichevan ASSR, Buzgov , leg. A. Ivanov ( ZISP) ; 1 ♂, 16.05.2014, Talysh, Yardymly vic., Deman , leg. V. Tikhonov ( OPB) . Kazakhstan. 1 ♀, 18.05.1899, Turgaisk Prov., Kaban-Kulak Mt. (20 verst SW Irgiz), leg. Sushkin ( ZISP) ; 1 ♀, 25.05.1907, Turgaisk Prov., Bolshie Barsuki, Chelkala lake , leg. Volman ( ZISP) ; 1 ♂, 29.06.1907, Uralsk , leg. B. Uvarov ( ZISP) ; 1 ♂, Karaganda Prov., 19.06.1958, 40 km S Zhana-Arka, Koksengir Mt. , leg. A. Zagulyaev ( ZISP) ; 1 ♂, 4– 5.05.2005, Akkum-Sagyz sands, leg. O. Novikov, slide OP 3141m ( OPB) ; 1 ♂, 19.06.2010, Kara- ganda Province, Zhaksy-Arganaty Mts., Karaatan , h= 500 m, 49 ° 14’ N 67 ° 01 E, leg. P. Gorbunov ( ZISP) ; 1 ♂, 30.06– 01.07.2010, 33 km N of Panfilov, Boro-Khoro Mts. , N 44 29.765 E 80 03.848, 1830 m, leg. S.K. Korb ( SKK) ; 2 ♂, 10.07.2011, Aktyubinsk Prov., Mugodzhary , N48° 32.172’ E58° 29.345’, 520 m, leg. S.K. Korb ( SKK) ; 11 ♂, 30.07.2011, Akmola Prov. near Zhaksy , N51° 52.769’ E67° 29.952’, leg. S.K. Korb ( SKK) ; 1 ♀, 26.04.2012, West Kazakhstan, Kumzhargan sands at Zhagabulak vill., 47 km SW of Emba town, 48º33’N, 57º37’E, leg. P. Gorbunov ( OPB) ; 1 ♀, 12.06.2012, Aktyubinsk Prov., Mugodzhary, Shauyryk river , 48°30’36.6” N 58°50’26.6” E, leg. Т. Trofimova and D. Shovkun ( ZISP) ; 1 ♂, 1 ♀, 30.06– 1.07.2012, Alma-Ata Prov., Panfilof distr., 25 km SW of Koktal, Ili river , leg. S. Rybalkin ( CKO) ; 1 ♂, 7– 8.05.2013, Alma-Ata Prov., 28 km NW of Bakanas, Ili river , leg. S.A. Knyazev ( CKO) ; 7 ♂, 3 ♀, 2.05.2014, Bol. Barsuki sands, 10 km E of Chelkar, 47°49’ N 59°46’ E, leg. P. Gorbunov ( ZISP) ; 2 ♂, 1 ♀, 1– 3.07.2015, Ili river valley near Kok-Tal bridge, N43°57.818’, E79°36.322’, 500 m, leg. S.K. Korb ( SKK) ; 2 ♂, 1 ♀, 4– 6.07.2015, Boro-Khoro Mts., Usek valley , N44° 28.082’ E79° 49.760’, 1260 m, leg. S.K. Korb ( SKK) ; 2 ♂, 07.07.2015, Boguty Mts., Chingilsu valley , N43° 34.825’ E78° 39.847’, 851 m, leg. S. K. Korb ( SKK) . Ukraine: 3 ♂, 17.05.1982, Kherson Prov., Chenromorsky Nature Reserve, Rybalchinsky Kordon , leg. I. Plyustsh, ex coll. A. Nekrasov, slide ffi1208 Nekrasov ( ZISP) . Russia. 4 ♂, 2 ♀, Narün , Henke (lectotype and paralectotype) ( ZMHU) ; 1 ♂, Sarepta , leg. unknown, ex coll. O. John ( ZISP) ; 4 ♀, Guberli , leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 25.06.[?], Guberli , leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 1 ♀, Spask , [leg. Eversmann], ex coll. Eversmann ( ZISP) ; 12 ♂, 4 ♀, 2.05.2010, Astra- khan Prov., Dosang , leg. S.K. Korb ( SKK) ; 1 ♂, 4.05.2010, Saratov Prov., Melovoe , leg. S.K. Korb ( SKK) ; 2 ♂, 7.05.2010, Volgograd Prov., Eruslan river valley, leg. S.K. Korb ( SKK) ; 1 ♂, 12.05.2019, Volgograd Prov., 17 km SSE Frolovo , 49°39’39”N 43°28’23”E, leg. A. Samus ( SKV) ; 1 ♂, 1 ♀, Caucasus , leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 1885, Caucasus , leg. Lederer. ex coll. Acad. Petrop ( ZISP) ; 1 ♀, 22.06.1893, Alupka , leg. Melioransky ( ZISP) ; 1 ♂, 2 ♀, 12.05.1904, 22– 29.06.1905, Ai-Petri , slope near Yalta, leg. S. Tshet- verikov ( ZISP) ; 1 ♂, 3.06.1905, St [ary] Kr [ym], [leg. Pliginsky] ( ZISP) ; 7 ♂, 4 ♀, 1.06.1905, 20.04– 22.05.1906, Agarmysh , [leg. Pliginsky], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 11.06.1905, distr. Jaltensis, loc. Oreanda , leg. S. Tshetverikov ( ZISP) ; 1 ♂, 8 ♀, 13.06.1905, 28.05.1907, 26.05– 26.06.1908, distr. Jaltensis, mons Ai-Petri , southern slope, leg. S. Tshetverikov, ex coll. S. Tshetverikov ( ZISP) ; 1 ♂, 26.05.1907, Simferopol , leg. G. and K. Khristoforov, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 3.06.1907, distr. Jaltensis , loc. Mis- chor, lower border of pine forest, leg. S. Tshetverikov ( ZISP) ; 2 ♂, 4 ♀, 26.05– 24.06.1908, distr. Jaltensis, loc. Koreiz, stony scree near Alupka , leg. S. Tshetverikov, ex coll. S. Tshetverikov ( ZISP) ; 1 ♂, 1 ♀, 4.06.1910, distr. Jaltensis, mnt. Ai-Petri , screes over Alupka, leg. S. Tshetverikov, ex coll. S. Tshetverikov ( ZISP) ; 1 ♂, 1 ♀, 23– 25.06.1913, distr. Theodosia , Staryi Crim, leg. S. Tschetverikov ( ZISP) ; 1 ♀, 31.07.1924, Karadag , leg. A. Dyako- nov ( ZISP) ; 2 ♂, 29.06.1926, 20.06.1944, Daghestan, Chodzhalmachi , leg. M. Rjabov ( ZISP) ; 1 ♀, 3.08.1926, Kumtorkale , leg. M. Ryabov ( ZISP) ; 1 ♀, 15.06.1992, Daghestan, Rutul , leg. O. Pekarsky, slide OP3140f ( OPB) ; 1 ♂, 27.07.2001, 25 km SW Buynaksk, Balakhani vill., leg. A. Magomedova ( ZISP) ; 2 ♂, 24.06.2002, Ay-Petri yayla, h=1300 м, leg. S. Sinev ( ZISP) ; 1 ♂, 18.06.2008, Krasnodar Prov., Seversky distr., Papay Mts. , arid semiforest, southern slope steppes, h=700–800 м, leg. V. Stshurov ( ZISP) ; 1 ♀, 28.05.2009, Crimea, Chiginitra , leg. V. Savchuk ( OPB) ; 1 ♂, 29.06.2010, Old Crimea, Agarmish Mt. , leg. V. Savchuk ( OPB) . Tajikistan. 1 ♂, 1 ♀, 16.07.2011, Shugnansky Mts., Gunt river valley, 2670 m, N 37 41.887, E 71 55.821, leg. A. Nikolaev, slide OP 3171m, OP3172f ( OPB) ; 1 ♂, 2 ♀, 17– 18.06.2012, Gissar Mts., distr. Varzob, vill. Kondara , 1150–1200 m, leg. E. Rutjan, slide OP 3171m, OP3172f, OP3182f ( OPB) . Turkey. 1 ♀, Asia m[inor], leg. unknown, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 26– 27.07.1984, Prov. Ağrý, 39º57´N, 43º14´E, Asgüneyi Dağlarý 2000 m, 3 km nö Çu- maçay, leg. H. Hacker ( HNHM) ; 1 ♂, 4.07.2002, Prov. Hakkari, 30 km NE of Hakkari, 1500 m, 37º41´N, 43º55´E, leg. B. Benedek and T. Csővári ( HNHM) ; 2 ♂, 1 ♀, 7– 8.07.2002, Prov. Agri, 2000 m, Karasu-Aras mts , 5 km SE of Sarican, 39º47´N, 42º28´E, leg. B. Benedek and T. Csővári ( HNHM) . Turkmenistan. 1 ♂, Askh [abad], [leg. Chris- toph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, Aschab [ad], ex coll. Bramson ( ZISP) ; 2 ♂, Tekke ( ZMHU) ; 1 ♂, 1 ♀, 29.05– 10.06.1882, Nochur , leg. Chr [istoph]. ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 2 ♀, 20.05.[?], Aidara , leg. Eylandt, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♀, 4.07.1892, Aidere , [leg. Christoph], ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 5 ♂, 6 ♀, 15.06– 20.08.1894, Transcaspien, Sumbar , leg. Herz, ex coll. Grand Duke Nikolai Mikhailovich ( ZISP) ; 1 ♂, 29.04.1952, 55 km E Kara-Kala, Ai- Dere , leg. V. Kusnetzov ( ZISP) ; 1 ♀, 27.06.1952, Kara-Kala , garden of VIR, leg. V. Kuznetsov ( ZISP) ; 5 ♂, 2 ♀, 7.04– 6.07.1953, 15 km NO Kara-Kala, Iol-Dere , leg. V. Kuznetsov ( ZISP) ; 5 ♂, 5 ♀, 18.06– 18.07.1954, 22.06.– 7.08.1956, mts. Kopet-Dag , valles Boboso, leg. W. Potopolskij ( ZISP) ; 1 ♂, 3 ♀, 7.07.1954, 16.05.1955, Aschabad circ., Tschuli, leg. W. Potopolskij ( ZISP) ; 1 ♂, 1 ♀, 12.06.1960, Askhabad vic., leg. Potopolsky ( ZISP) ; 1 ♂, 3 ♀, 1.05– 11.06.1986, Aidere , h=850 м, leg. M. Falkovich ( ZISP) . Iran. 1 ♀, 31.05– 2.06.2014, Prov. Mazandaran, near Gaschar vill., N 36º6’36.48”, E 51º20’1.85”, leg. D. Kasatkin, I. Shokhin, slide OP3182f ( OPB) .

Figures

FIGURES 1–21. Drasteria specimens. 1–3. D. cailino obscura (Staudinger, 1901), lectotype (1: upperside; 2: underside; 3: labels). 4–12. D. cailino baigakumensis (John, 1917), lectotype (4: upperside; 5: underside; 6: labels), Almaty vicinity, Kazakhstan (7), paralectotypes (8, 9), Tekes, Turkmenistan (10); Bishkek, Kyrgyzstan (11); Padkhinov vill., Tajikistan (12). 13–17. D. sculpta sculpta (Püngeler, 1904), lectotype (13: upperside; 14: underside; 15: labels), Almaty, Kazakhstan (16), paralectotype (17). 18–21. D. sculpta pamira (John, 1921), Pakistan (18), lectotype (19: upperside; 20: underside; 21: labels).

FIGURES 297–304. Drasteria habitats. 297. 25 km N of Panfilov, Dzhungarian Alatau, Kazakhstan (one of the authors in the collecting trip 2010). 298. Ili river valley near Koktal bridge, Almaty Prov., Kazakhstan (Russian Entomological Society and Institute of Zoology of Kazakhstan collecting trip, 2010). 299. Sands near Dosang environs, Astrakhan Prov., Russia. 300. Saksaul forest near Bakanas, Almaty Prov., Kazakhstan. 301. Chulak gorge, foothills of Dzhungarian Alatau Mts. in the Altyn-Emel Nature Reserve, Kazakhstan. 302. Ili river shore near Singing Dunes, Altyn-Emel Nature Reserve, Kazakhstan. 303. Katutau mountains, Altyn-Emel Nature Reserve, Kazakhstan. 304. Koram environs, northern slopes of Transilian Alatau Mts., Kazakhstan. Photos: S.K. Korb.