Forstercooperia ulanshirehensis, Wang & Bai & Meng & Wang, 2018

Wang, Hai-Bing, Bai, Bin, Meng, Jin & Wang, Yuan-Qing, 2018, A New Species of Forstercooperia (Perissodactyla: Paraceratheriidae) from Northern China with a Systematic Revision of Forstercooperiines, American Museum Novitates 2018 (3897), pp. 1-44 : 5-6

Forstercooperia ulanshirehensis , sp. nov.

HOLOTYPE: A nearly complete cranium (AMNH 21608) ( figs. 2–4 ).

REFERRED SPECIMENS: A pair of complete mandibles ( IVPP V20155 ) ( figs. 5 , 6 ), a fragmentary maxilla with P3–M3 ( IVPP V20154 ) ( figs. 7A, 7B ); a fragmentary mandible with the lower canine and p1–4 ( IVPP V20156 ) ( figs. 7C, 7D ), and a fragmentary mandible with p3–m3 ( AMNH 22101) ( fig. 8 ).

ETYMOLOGY: The specific name ulanshirehensis refers to the Ulan Shireh Formation, in which AMNH 21608 was collected.

HORIZON AND LOCALITY: AMNH 21608 (field number, 613), Ulan Shireh Formation, Wulanhuxiu site (previously known as 8 miles north of Tukhum Lamasery ) of Erlian Basin, Nei Mongol, China ; AMNH 22101 (field number, 602), Ulan Shireh Formation, Wulantaolegai site (previously known as 4 miles north of Tukhum ) of Erlian Basin, Nei Mongol, China ; IVPP V20154 , V20155 , and V20156 , Irdin Manha Formation , Irdin Manha site, Erlian Basin, Nei Mongol, China . Middle Eocene.

DIAGNOSIS: Small species of Forstercooperia , the length of upper premolar series about 81 mm, about 25% smaller than that of Forstercoopeia totadentata (~ 108 mm), about 20% smaller than that of Forstercooperia mazhuangensis (~ 103 mm) (see appendix, tables 2–3). Upper premolars transversely slightly narrower. Protocone and hypocone of P2 barely separate, metaloph of P2 narrow and posterolingually oriented. Metalophs (metaconules) of P3–4 short. Entoconid of p4 present.

Figures

FIG. 2. Cranium of F. ulanshirehensis (AMNH 21608) in lateral view. Abbreviations: for, foramen orbitale; fr, frontal; fro, foramen rotundum; iof, infraorbital foramen; ju, jugal; lc, lacrimal; mr, median ridge of postglenoid process; mx, maxilla; na, nasal; oc, occipital condyle; opf, optic foramen; pa, parietal; pf, preorbital fossa; pgp, postglenoid process; pop, postorbital process; pocp, paroccipital process; ppf, posttympanic-paroccipital foramen; ptp, posttympanic process; sc, sagittal crest; sq, squamosal; tc, temporal crest.

FIG. 3. Cranium of F. ulanshirehensis (AMNH 21608) in dorsal view. Abbreviations: fr, frontal; iof, infraorbital foramen; ju, jugal; mx, maxilla; na, nasal; nf, nutrient foramen; pa, parietal; sc, sagittal crest; sq, squamosal.

FIG. 4. Cranium of F. ulanshirehensis (AMNH 21608) in ventral view. Abbreviations: bo, basioccipital; bs, basisphenoid; gch, partial great cornu of hyoid; hf, hypoglossal foramen; ju, jugal; mr, median ridge of postglenoid process; mx, maxilla; oc, occipital condyle; of, oval foramen; pac, posterior opening of the alisphenoid canal; pal, palatine; pgp, postglenoid process; pocp, paroccipital process; ppa, pterygoid process of alisphenoid; ptp, posttympanic process.

FIG. 5. Mandibles of F. ulanshirehensis (IVPP V20155) in occlusal view. Abbreviations: cdp, condylar process; crp, coronoid process; pcp, postcotyloid process.

FIG. 6. Mandibles of F. ulanshirehensis (IVPP V20155) in lateral view. Abbreviations: cdp, condylar process; crp, coronoid process; mf, mandibular foramen; pcp, postcotyloid process.

FIG. 7. Dentition of F. ulanshirehensis in lateral and occlusal view. A, IVPP V20154 in lateral view; B, IVPP V20154 in occlusal view; C, IVPP V20156 in occlusal view; D, IVPP V20156 in lateral view.

FIG. 8. Dentition of F. ulanshirehensis in occlusal and lateral view (AMNH 22101). A, occlusal view; B, lateral view.