Ochthephilus andalusiacus ( Fagel, 1957 )

Makranczy, György, 2014, Revision of the genus Ochthephilus Mulsant & Rey, 1856 (Coleoptera: Staphylinidae, Oxytelinae), Revue suisse de Zoologie 121 (4), pp. 457-694 : 557-560

Ochthephilus andalusiacus ( Fagel, 1957) Figs 309-313, 332, 360, 364, 540, 581

Ancyrophorus andalusiacus Fagel, 1957: 704 . – Fagel, 1968: 192.

Ochthephilus andalusiacus (Fagel) . – Herman, 1970: 384. – Pope, 1977: 24. – Makranczy, 2001: 180. – Lott, 2008: 18.

Ancyrophorus rivularis Smetana, 1967: 384 , 388.

TYPE MATERIAL EXAMINED: Ancyrophorus andalusiacus – HOLOTYPE (3): “ Andalousie , environ de Ronda, Puente de la Ventilla, route de Malaga, km 7 [*+36.73/-05.08*], 8-20.V.1956, leg. G. Fagel ” ( ISNB) . – PARATYPES (2): same data as holotype ( ISNB, 1, BMNH, 1). Ancyrophorus rivularis – HOLOTYPE (3): “ Slovakia, Vysoké Tatry , Tatranská Lomnica [*+49.16/+20.28*], 1.VII.1956, leg. [A.] Smetana” (coll. Smetana, CNCI) . – PARATYPE (1): “ Slovakia, Lukov - Bardejov , Laborec (stream) [*+49.30/+21.25*], 7.VII.1959, leg. [A.] Smetana” (coll. Smetana, CNCI, 1) .

OTHER MATERIAL: see Appendix.

REDESCRIPTION: Forebody as in Fig. 540. Measurements (n=10): HW = 0.53 (0.50-0.56); TW = 0.48 (0.45-0.50); PW = 0.58 (0.53-0.62); SW = 0.70 (0.65-0.78); AW = 0.76 (0.71-0.80); HL = 0.39 (0.34-0.43); EL = 0.19 (0.18-0.20); TL = 0.06 (0.05- 0.07); PL = 0.45 (0.42-0.49); SL = 0.92 (0.85-0.98); SC = 0.84 (0.77-0.90); FB = 1.79 (1.65-1.89); BL = 3.19 (3.00-3.44) mm. Head, pronotum and abdomen blackish dark brown. Elytra reddish dark to medium brown, scutellar area to shoulders darker, blackish; occasional specimens with lighter elytra, even with lighter spot behind middle of suture. Antennae and mouthparts dark brown with occasional reddish tint, legs reddish medium to dark brown. Body with greasy lustre mostly due to elytral setation plus forebody punctation and microsculpture. Pubescence rather fine and moderately dense, shorter and stronger (regularly spaced) on elytra, abdominal tergites with finer and longer setae, especially adjacent to laterosternites. Head anteriad eyes and near inner posterior margin of eye with stronger and darker bristles, as well as pronotal margin and middle of tibiae. Elytral apex with one or two slightly larger setae near sutural corners. Last tarsomere with a few setae only.

Forebody. Antenna as in Fig. 581, antennomere 6 shorter and less wide than neighbours (articles 5 and 7). Clypeus almost impunctate (colliculate microsculptured), trapezoid, corners rounded, anterior edge gently arched; separated by impressed transversal line (frontoclypeal suture) across a shinier area. Supraantennal prominences well developed, feebly separated from clypeus/vertex by impressions. Vertex with oblique impressions in middle almost joining in V-shape. Temples bulging, evenly curved, little shorter than half of eye length. Neck separated by an impressed transversal groove, microsculpture much stronger than on head, with transverse cells, no setation. Pronotum with a narrow marginal bead, visible to anterior pronotal corners. Posterior pronotal angles well-formed, just slightly obtuse-angled, sides in posterior 1/3 very gently concave. 'Anchor' fully formed, longitudinal midline as a slightly elevated, impunctate, weakly microsculptured line, parallel to this line two gentle, semilongitudinal elongate elevations in anterior half of disc. In corners of anchor feeble, oblique impressions directed outwards, in middle at sides of midline two smaller impressions. Elytra slightly broadening posteriorly, sutural corners narrowly rounded; apical sides slightly oblique and in inner halves gently convex. Elytral surface rather even with two shallow, very elongate impressions behind scutellum. Head with fine coriaceous/colliculate microsculpture, fading on elevated parts, stronger in impressions, on pronotum microsculpture slightly stronger and more even. Punctation on head sparse, mostly confined to posterior part and sides, on pronotum more evenly spaced, average interspaces much larger than puncture diameters; elytral punctation more even and regularly spaced, average interspaces (with indistinct coriaceous microsculpture) about as puncture diameters, punctures discrete.

Abdomen. Compared to forebody, abdomen with much more sparse, finer, less distinct punctation, microsculpture on tergal apices fine coriaceous with moderately transverse cells. Tergite VII posterior margin with palisade fringe unmodified in middle (nearly uniform breadth). Tergite VIII (Fig. 360) basal edge evenly arched, with small concavity in middle of basal sclerotized band; apical edge with sinuate (protruding) corners, and broad, moderately deep emargination in between. Sternite VIII with rounded apical corners, apex in males shallowly concave laterally, gently sinuate

FIGS 300-303

(300-301) Ochthephilus qingyianus sp. n.; head and pronotum (300), elytra (301). (302-303) O. szeli sp. n.; head and pronotum (302), elytra (303). All SEM, dorsal views. Scale bar = 0.44 mm for 302, 0.5 mm for 300, 303, 0.55 mm for 301.

in middle; in females slightly more sinuate (convex) in middle. Tergite X with modified apical margin, in males as in Fig. 332, in females as in Fig. 364. Aedeagus as in Fig. 309, inner sclerites as in Figs 310-311. Female ringstructures as in Figs 312-313.

COMPARATIVE NOTES: The triad of species where this taxon can easily be confused also includes O. omalinus and O. venustulus , sharing the same formation of mid-antennal articles but differing in the apices of tergite X in both sexes: not irregularly serrate in the latter two species. Colouration can be confusing, as all three species may have lighter elytra, even with a distinct light spot around the suture behind the middle. In most cases O. andalusiacus has to be separated from O. omalinus : the latter has the middle apex of tergite almost straight but with a little concavity in the middle; O. andalusiacus has a peculiar formation of the apical corners (strongly sclerotized and evenly arched on inside), while the middle apex is just perceptibly convex, with no concavity in the centre. The unusual specimens with labels "Reinfeld i. Holst., 14.IV.1926, Bernitz" were identified by G. Benick as ' O. strandi ' and such specimens are responsible for the false record of the species from Central Europe.

DISTRIBUTION: Known from North Africa, Europe, Turkey and apparently introduced to eastern Canada (coastal areas).

BIONOMICS: Specimens were collected from riverbank and seashore (vegetable) debris, at waterfall, sandy-gravelly bank and debris on wet mosses, mainly willow moss (Fontinalis sp.), also captured by car-net.