Digitonthophagus catta

Génier, François & Moretto, Philippe, 2017, Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 4248 (1), pp. 1-110 : 24

Digitonthophagus catta species group

Diagnosis. Pronotal punctures scattered and small on posterior half ( Figs. 58 , 93); surface behind anterior pronotal angles concave; male with two erected cephalic horns ( Figs. 30 , 32 ); females with anterior pronotal tubercles well developed and deeply obliquely concave (Figs. 94, 96); subapicodorsal lobe of FLP sclerite enlarged and sclerotized ( Figs. 144 , 147 ); FLP left surface with a conspicuous apophysis ( Figs. 144 , 147 ).

Figures

FIGURES 55 – 62. Male pronotum, dorsal view. 55. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 56. D. uks, holotype; 57. D. sahelicus, holotype; 58. D. catta, alloreferent (Tamil Nadu State, India); 59. D. eucatta, holotype; 60. D. falciger, holotype; 61. D. biflagellatus, holotype; 62. D. gazella, non type (Central Province, Zambia).

FIGURES 23 – 30. 23, 25, 27, 29. Male head, dorsal view; 24, 26, 28, 30. Male head, frontal view. 23 – 24. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 25 – 26. D. uks, holotype; 27 – 28. D. sahelicus, holotype; 29 – 30. D. catta, alloreferent (Tamil Nadu State, India).

FIGURES 31 – 38. 31, 33, 35, 37. Male head, dorsal view; 32, 34, 36, 38. Male head, frontal view. 31 – 32. Digitonthophagus eucatta, alloreferent; 33 – 34. D. falciger, holotype; 35 – 36. D. biflagellatus, holotype; 37 – 38. D. gazella, non type (Central Province, Zambia).

FIGURES 135 – 146. Internal sac sclerites (FLP, A + SA). 135, 138, 141, 144. Dorsal view; 136, 139, 142, 145. Ventral view. 137, 140, 143, 146. Right lateral view. 135 – 137. Digitonthophagus bonasus, alloreferent (Tamil Nadu State, India); 138 – 140. D. uks, holotype; 141 – 143. D. sahelicus, holotype; 144 – 146. D. catta, alloreferent.

FIGURES 147 – 158. Internal sac sclerites (FLP, A + SA). 147, 150, 153, 156. Dorsal view; 148, 151, 154, 157. Ventral view. 149, 152, 155, 158. Right lateral view. 147 – 149. Digitonthophagus eucatta, alloreferent (Tamil Nadu State, India); 150 – 152. D. falciger, holotype; 153 – 155. D. biflagellatus, holotype; 156 – 158. D. gazella, non type (Antisiranana Province, Madagascar).