Nanophareus Roewer, 1929

Hara, Marcos Ryotaro, Pinto-Da-Rocha, Ricardo & Kury, Adriano Brilhante, 2012, Revision of Nanophareus, a mysterious harvestman genus from Chile, with descriptions of three new species (Opiliones: Laniatores: Gonyleptidae), Zootaxa 3579, pp. 37-66 : 46

Nanophareus Roewer, 1929 , new subfamilial assignment

Nanophareus Roewer 1929: 280 (key, desc); Canals 1936: 69 (cat); Cekalovic 1985: 24 (cat); Pinto-da-Rocha 1997: 166 (syst); Kury 2003: 105 (cat), 259 (syst).

Type species: Nanophareus palpalis Roewer, 1929 , by monotypy.

Diagnosis. Easily distinguished from other Pachylinae by: Pedipalpal patella inserted dorsally on tibia; pedipalpal tibia basal ventrally curved 90 degrees in lateral view, with retrolateral distal or subdistal socket setae longest and bifid apically (one short and one long setae); pedipalpal patella and tibia with dorsal tubercles (except N. bosqenublado sp. nov.); a widened and low ocularium that superficially appears divided (except N. araucanus sp. nov.).

Redescription. Male: Dorsum: Anterior margin of carapace with a median frontal hump and a row of tubercles on each side. Ocularium widened (approximately ½ of carapace width, rendering a divided appearance) or not (approximately ½ of carapace width). Ocularium unarmed, with sparse tubercles, or with median pair of spines. Dorsal scutum with five transverse grooves delimiting four scutal areas. Scutal area I divided in half by a longitudinal median groove; III unarmed ( N. bosqenublado sp. nov., N. bipartitus sp. nov. and N. palpalis ) or with a pair of spines ( N. araucanus sp. nov.). Body shape roughly pyriform, wider between scutal grooves I and IV. Lateral margin of dorsal scutum with an external row of enlarged tubercles inserted among small ones (innapplicable in N. bosqenublado because it has no external row of tubercles along the lateral margin of the dorsal scutum). Posterior margin of dorsal scutum and free tergites I–III each with a row of tubercles.

Chelicera: Similarly shaped in males and females. Segment I with a weakly marked bulla, smooth or with sparse tubercles; both fingers toothed.

Pedipalpus: Coxa dorsobasally inflated, trochanter inflated with sparse tubercles. Femur unarmed, without prolateral subapical seta. Patella dorsally inserted in tibia. Tibia basal ventrally curved 90 degrees in lateral view, with variable setation, retrolateral distal or subdistal socket setae longest and bifid apically (1 short and 1 long setae). Tarsus with variable setation.

Legs: Coxa I with 1 prolateral, 1 retrolateral apophyses; II idem, with a small dorsomedian tubercle (absent in N. bosqenublado ), 1 retrolateral apophysis fused at apex with prolateral apophysis of coxa III; III with 1 prolateral, 1 retrolateral apophyses; IV irregularly tuberculate, with 1 prolateral apical apophysis more developed in male than female. Coxa IV surpassing dorsal scutum in dorsal view, being visible in all extension. Trochanter IV armed prolaterally. Femora–tibiae I–IV with tubercles approximately arranged in six longitudinal rows (a pro- and a retrodorsal row, a pro- and a retroventral row, a pro- and a retrolateral row). Femur I–III roughly straight and unarmed; IV roughly straight or strongly sigmoid with variable armature.

Penis: Glans with stylus and ventral process, without dorsal process. Ventral process of glans with blunt apex or apex with lateral projections. Ventral plate developed, almost rectangular (widened at basal half), anterior margin straight, distal half slightly thickened, with 3 pairs of distal conical setae, 1 pair of median small setae, 3 pairs of basal setae.

Female: Similar to male, coxa IV shorter than in male (reaching between scutal grooves III and IV), with shortened conical, pointed prolateral apical apophysis (its length close to its basal diameter). Trochanter IV unarmed prolaterally, femora–metatarsi I–IV unarmed.