Hemirhagerrhis Boettger, 1896

Marques, Mariana P., Ceríaco, Luis M. P., Blackburn, David C. & Bauer, Aaron M., 2018, Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola Atlas of Historical and Bibliographic Records (1840 - 2017), Proceedings of the California Academy of Sciences 65, pp. 1-501 : 333-334

Genus Hemirhagerrhis Boettger, 1896

Hemirhagerrhis viperina (Bocage, 1873) WESTERN BARK SNAKE Psammophylax viperinus Bocage 1873a:222 . Holotype: MBL 1715 (collector J.A. d’Anchieta), destroyed by fire 18 March 1978. Type locality: “Dombe” [= Dombe Grande], Benguela Province, Angola. Psammophylax nototaenia : Bocage (1895a:109). Amplorhinus nototaenia : Boulenger (1915:211). Amplorhinus nototaenia ( Psammophylax nototaenia ): Monard (1937b:128). Hemirhagerrhis nototaenia viperinus : Bogert (1940:76), FitzSimons (1962:208), Laurent (1964a:112),

Broadley (1990:120). Hemirhagerrhis viperina : Broadley (1997c:164), Branch (1998:86), Broadley and Hughes (2000:7), Wallach et al. (2014:321). Hemirhagerrhis nototaenia : Branch (1998:86), Wallach et al. (2014:320).

Global conservation status (IUCN): Not Evaluated.

Global distribution: The species is known from southwestern Angola to northern Namibia.

Ocurrences in Angola (Map 292): The species occurs in the southwestern Angola. Huambo: “Huambo” [-12.83333, 15.66667] ( Bogert 1940:75; Broadley 1997c:164; Broadley and Hughes 2000:9). Benguela: “Dombe” [-12.95000, 13.10000] ( Bocage 1873a:222, 1895a:109; Monard 1937b:128; FitzSimons 1962:208; Broadley 1990:120, 1997c:161, 164; Broadley and Hughes 2000:9); “Hanha” [-12.25000, 13.75000] MAP 292. Distribution of Hemirhagerrhis viperina in

Angola.

( Bogert 1940:76; Broadley 1997c:164; Broadley and Hughes 2000:9). Huíla: “Humpata, Fazenda Bumbo” [-15.20000, 13.00000] ( Laurent 1964a:112; Broadley 1997c:164; Broadley and Hughes 2000:9); “ 8 km from Tundavala to Sa da Bandeira” [-15.83333, 13.40000] ( Broadley 1997c:164; Broadley and Hughes 2000:9); “ 5 km S of Chibemba” [-15.75000, 14.08333] ( Broadley 1997c:164; Broadley and Hughes 2000:9). Namibe: “Huxe” [-12.71667, 13.20000] ( Broadley 1997c:164; Broadley and Hughes 2000:9); “Lungo” [-14.31667, 13.20000] ( Broadley 1997c:164; Broadley and Hughes 2000:9); “Munhino” [-14.91667, 13.00000] ( Bogert 1940:76; Broadley 1997c:164; Broadley and Hughes 2000:9); “Maconjo” [-15.01667, 13.20000] ( Bocage 1895a:109; Broadley 1997c:161, 164; Broadley and Hughes 2000:9); “Caraculo” [-15.01667, 12.66667] ( Broadley 1997c:164; Broadley and Hughes 2000:9); “Capangombe” [-15.10000, 13.15000] ( Bocage 1895a:109; Broadley 1997c:161, 164; Broadley and Hughes 2000:9). Cunene: “Humbe” [-16.68333, 14.90000] ( Bocage 1895a:109; Broadley 1997c:161).

Taxonomic and distributional notes: Bocage (1873a) described Psammophylax viperinus based on one individual from “Dombe” collected by Anchieta in Benguela Province. Subsequently he ( Bocage 1895a) synonymized P. viperinus with P. nototaenia ( Günther, 1864) and recorded six specimens from southwestern Angola: two from “Maconjo,” one from “Capangombe,” two from “Humbe,” and the “Dombe ” viperinus speciemen. Broadley (1997c) examined the material in Museu Bocage in 1968, and found only five specimens, the one from “Dombe” and two each from “Maconjo” and “Capangombe” (the reason for the discrepancy in the number of specimens from Capangombe is unknown). He suggested that the missing specimens from “Humbe” may have been typical H. nototaenia . However, H. nototaenia extends from Kenya to South Africa and Mozambique, whereas H. viperina is widely distributed from southwestern Angola to northern Namibia ( Broadley 1990, 1997c; Broadley and Hughes 2000), suggesting that all records from Angola are correctly interpreted as H. viperina .